Additional to the Fungi: FUNGI IMPERFECTI. Incompletely known
(_Saccharomyces_, _Oidium_-forms, etc.).
Class 1. =Phycomycetes (Algal-Fungi).=[12]
This group resembles _Vaucheria_ and the other Siphoneæ among the
Algæ.
ORGANS OF NUTRITION. The mycelium is formed of a single cell, often
thread-like and abundantly branched (Fig. 78). Vegetative propagation
by chlamydospores and oidia. Asexual reproduction by endospores
(sometimes _swarmspores_) and conidia. Sexual reproduction by
conjugation of two hyphæ as in the Conjugatæ, or by fertilisation of an
egg-cell in an oogonium. On this account the class of the Phycomycetes
is divided into two sub-classes: ZYGOMYCETES and OOMYCETES.
Sub-Class I. =Zygomycetes.=
Sexual reproduction takes place by zygospores, which function as
resting-spores, and arise in consequence of _conjugation_ (Fig. 81); in
the majority of species these are rarely found, and only under special
conditions. The most common method of reproduction is by endospores, by
acrogenous conidia, by chlamydospores, or by oidia. _Swarmspores are
wanting._ Parasites and saprophytes (order 6 and 7). The zygospores are
generally produced when the formation of sporangia has ceased; _e.g._
by the suppression of the sporangial-hyphæ (_Mucor mucedo_), or by
the diminution of oxygen; _Pilobolus crystallinus_ forms zygospores,
when the sporangia are infected with saprophytic _Piptocephalis_ or
_Pleotrachelus_.
=A.= Asexual reproduction only by sporangia.
Order 1. =Mucoraceæ.= The spherical sporangia contain many spores.
The zygospore is formed between two unicellular branches (gametes).
The unicellular mycelium (Fig. 78) of the Mucoraceæ branches
abundantly, and lives, generally, as a saprophyte on all sorts of
dead organic remains. Some of these Fungi are known to be capable of
producing _alcoholic fermentation_, in common with the Saccharomyces.
This applies especially to _Chlamydomucor racemosus_ (_Mucor
racemosus_), when grown in a saccharine solution, and deprived of
oxygen; the mycelium, under such conditions, becomes divided by
transverse walls into a large number of small cells. Many of these
swell out into spherical or club-shaped cells, and when detached from
one another become chlamydospores, which abstrict new cells of similar
nature (Fig. 79). These chlamydospores were formerly erroneously
termed “mucor-yeast,” but they must not be confounded with the
yeast-conidia (page 94). They are shortened hyphæ, and are not conidia
of definite size, shape, and point of budding. Oidia are also found in
_Chlamydomucor_.
[Illustration: FIG. 78.--_Mucor mucedo._ A mycelium which has
sprung from one spore, whose position is marked by the *: _a_,
_b_, _c_ are three sporangia in different stages of development;
_a_ is the youngest one, as yet only a short, thick, erect
branch; _b_ is commencing to form a sporangium which is larger in
_c_, but not yet separated from its stalk.]
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