Genera: _Saprolegnia_, whose swarmspores disperse immediately
after having left the sporangium. _S. ferax_ is the
cause of a disease in fish (“Salmon disease”) and in the
crayfish.--_Achlya_, whose swarmspores accumulate in a hollow
ball before the mouth of the sporangium.--_Leptomitus_ has
strongly indented hyphæ, causing a “linked” appearance.
_L. lacteus_ is frequent in the waste matter from sugar
factories.--_Monoblepharis_ deviates from the others by the
greater development of its fertilising process; the oosphere,
situated in an open oogonium, becoming fertilised by self-motile
spermatozoids, which are provided with a cilium at the posterior
end.
Class 2. =Mesomycetes.=
The Mesomycetes are intermediate forms between the Phycomycetes and
the Higher Fungi. In the vegetative organs, and in the multicellular
hyphæ, they resemble the Higher Fungi; the methods of reproduction,
however, show the characters of the Phycomycetes, namely sporangia
and conidiophores of varying size and with varying number of spores;
definite and typically formed asci and basidia are not present. Sexual
reproduction is wanting. The HEMIASCI are transitional between the
Phycomycetes and the Ascomycetes, the HEMIBASIDII (Brand-Fungi) form
the transition to the Basidiomycetes.
Sub-Class 1. =Hemiasci.=
The Hemiasci are Fungi with _sporangia_ which, _although resembling
asci_, yet have _not_, however, _a definite form and a definite number
of spores_. Besides endospores, conidia, chlamydospores and oidia are
found.
Order 1. =Ascoideaceæ.= _Ascoidea rubescens_ forms irregular,
reddish-brown masses in the sap issuing from felled Beeches. It
has _free sporangia_, which resemble asci in their structure, in
the development and ejection, and in the definite shape and size
of the spores. The formation of the sporangia takes place when
the nutriment is nearly exhausted, and resembles that of the
conidia, since they are developed from the end of a hypha which
enlarges, and the swelling becomes separated by a transverse
wall. Within the sporangia numerous spores of a cap-like form
are developed, which are set free through an opening at the
apex. Sporangia are formed successively at the apex of the
same hypha, the second commencing to develope as the first is
dehiscing. Conidia and sporangia are not formed simultaneously;
the former may be considered as closed sporangia.
Order 2. =Protomycetaceæ.= _Protomyces pachydermus_ causes
hard swellings on the stems and leaf-stalks of the Cichorieæ
(_Taraxacum_, etc.). These swellings consist of _chlamydospores_
(resting-spores), which germinate and become free, ascus-like
sporangia, with numerous small spores. In nutritive solutions
the chlamydospores form conidia with yeast-like buddings. _P.
macrosporus_ on _Ægopodium_, and other Umbelliferæ.
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