A history of land mammals in the western hemisphereScott, William Berryman
History
A history of land mammals in the western hemisphere
Scott, William Berryman
Mammals; Mammals, Fossil
This family was abundantly represented in Europe from the middle Eocene
through the Oligocene, in Asia persisting even into the Pliocene, and
were abundant in the Oligocene of Egypt. Migrants from the Old World
reached America in White River times, but speedily died out, as they
did not survive into the upper Oligocene. The most fully known of these
animals is an American species of a European genus _†Bothriodon_. Almost
complete skeletons of this genus have been obtained in the channel
sandstones of the upper White River substage. In size and proportions,
_†Bothriodon_ was not unlike a domestic pig, but had a very long head
with slender, pointed snout; it had also a short neck, long body, short
limbs and feet. The primitive character of this genus is made clear
by many features of its structure; the molar teeth were extremely
low-crowned and their cusps were so imperfectly crescentic in form as to
be called _buno-selenodont_, as indicating their transitional nature,
and the upper molars had five cusps instead of four, a very primitive
feature. Another very significant character was the five-toed manus; the
first digit, or pollex, was much smaller than the others.
[Illustration: FIG. 196.—_†Bothriodon brachyrhynchus_, upper White River
stage. Restored from a skeleton in the museum of Princeton University.]
The second genus of the family which had American representatives was
_†Anthracotherium_, which was much like _†Bothriodon_, but even more
archaic in character; the molars could hardly be called selenodont at
all.
4. _†Oreodontidæ. †Oreodonts_
This was one of the most characteristic of North American artiodactyl
families, and its members were exceedingly abundant throughout the upper
Eocene, the whole Oligocene and Miocene, ending their long career in
the lower Pliocene. In distribution the family was exclusively North
American, and no trace of it has been found in any other continent.
In the course of their long history the †oreodonts underwent many
transformations and branched out into several distinct phyla, yet
through all these changes they remained singularly conservative, for the
transformations, some of them sufficiently bizarre, affected chiefly the
teeth and skull, the remainder of the skeleton changing but little. The
†oreodonts were all small or of moderate size, none of them surpassing
the Wild Boar in stature, nor was there any decided increase in size from
stage to stage. One and all, they were strange beasts. Dr. Leidy, who
first described and named most of the genera, spoke of them as combining
the characters of camel, deer and pig, and called them “ruminating hogs,”
a conception expressed in the names which he gave to some of them, such
as _†Merychyus_ and _†Merycochœrus_, both of which mean ruminant swine.
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