From the time of Nintoku (A.D. 313-399) until the introduction of Buddhism (A.D. 552), there were four uji whose chiefs participated conspicuously in the government of the country. The first was that of Heguri. It belonged to the Imperial class (Kwobetsu) and was descended from the celebrated Takenouchi-no-Sukune. In the days of the Emperor Muretsu (499-506), the chief of this uji attempted to usurp the throne and was crushed. The second was the Otomo. This uji belonged to the Kami class (Shimbetsu) and had for ancestor Michi no Omi, the most distinguished general in the service of the first Emperor Jimmu. The chiefs of the Otomo-uji filled the post of general from age to age, and its members guarded the palace gates. During the reign of Yuryaku the office of o-muraji was bestowed upon Moroya, then chief of this uji, and the influence he wielded may be inferred from the language of an Imperial rescript where it is said that "the tami-be of the o-muraji fill the country." His son, Kanamura, succeeded him. By his sword the rebellion of Heguri no Matori was quelled, and by his advice Keitai was called to the Throne. He served also under Ankan, Senkwa, and Kimmei, but the miscarriage of Japan's relations with Korea was attributed to him, and the title of o-muraji was not conferred on any of his descendants. The uji of Mononobe next calls for notice. "Monono-be" literally signifies, when expanded, a group (be) of soldiers (tsuwamono). In later times a warrior in Japan was called mono-no-fu (or bushi), which is written with the ideographs mono-be. This uji also belonged to the Kami class, and its progenitor was Umashimade, who surrendered Yamato to Jimmu on the ground of consanguinity. Thenceforth the members of the uji formed the Imperial guards (uchi-tsu-mononobe) and its chiefs commanded them. Among all the uji of the Kami class the Mononobe and the Otomo ranked first, and after the latter's failure in connexion with Korea, the Mononobe stood alone. During the reign of Yuryaku, the uji's chief became o-muraji, as did his grandson, Okoshi, and the latter's son, Moriya, was destroyed by the o-omi, Soga no Umako, in the tumult on the accession of Sushun (A.D. 588). The fourth of the great uji was the Soga, descended from Takenouchi-no-Sukune. After the ruin of the Heguri, this uji stood at the head of all the Imperial class. In the reign of Senkwa (536-539), Iname, chief of the Soga, was appointed o-omi, and his son, Umako, who held the same rank, occupies an important place in connexion with the introduction of Buddhism. It will be observed that among these four uji, Heguri and Soga served as civil officials and Otomo and Mononobe as military.