A Monograph on the Sub-class Cirripedia (Volume 1 of 2): The Lepadidae; Or, Pedunculated CirripedesDarwin, Charles
Science
A Monograph on the Sub-class Cirripedia (Volume 1 of 2): The Lepadidae; Or, Pedunculated Cirripedes
Darwin, Charles
Cirripedia; Lepadidae
_Scuta._--In three of the species the two segments, named the occludent
and basal, appear like separate valves, but these, by dissection, can be
most distinctly seen to be united at the rostral angle. The primordial
valve, formed of the usual hexagonal tissue, is elliptic, elongated, and
placed in the direction of the occludent segment; calcification
commences at its upper point, so as to form the occludent segment, and
afterwards at its lower point, but rectangularly outwards, to form the
basal segment; in the minute space between these two points of the
primordial valve, there is, in four of the species, no calcification; so
that the two segments are united by what may be called a flexible hinge;
in _D. orthogonia_ the two calcareous segments are absolutely
continuous. The occludent segment is longer than the basal segment; it
either runs close along the orifice, or in the upper part bends inwards;
both segments are narrow, except in _D. Warwickii_, in which the basal
segment is moderately broad; the two segments are placed at an angle,
varying from 45° to 90°, to each other. The capitulum generally extends
for a little space beneath the basal segments of the scuta, where it
contracts to form the peduncle.
The _Terga_ present singular differences in shape, and are described
under the head of each species; scarcely any point can be predicated of
them in common, except that they are flat and thin.
The _Carina_ is much bowed, narrow, and internally either slightly
concave or convex and solid; the upper end extends far up between the
terga; the lower end is formed by a rectangularly inflected, imbedded,
triangular or oblong disc, deeply notched at the end, or as in _H.
Lowei_, of a fork, the base, however, of which is wider than the rest of
the carina, so as to present some traces of the disc-like structure of
the other two species; or lastly, as in _D. orthogonia_, it terminates
in a crescent-formed cup.
_Peduncle._--This is narrow, compressed, and about as long, or twice as
long, as the capitulum; in _D. Warwickii_ it is studded with minute
beads of yellowish chitine.
_Size._--Small, with a capitulum scarcely exceeding a quarter of a inch
in length.
_Filamentary Appendages._--None. There are two small ovigerous fræna,
which, in _D. Warwickii_, had the glands collected in seven or eight
little groups on their margins.
_Mouth._--Labrum highly bullate, with small teeth on the crest; palpi
small, not thickly covered with spines. _Mandibles_ narrow, with three
or four teeth. Maxillæ small, with a notch beneath the two or three
great upper spines; lower part bearing only a few pair of spines,
generally not projecting, but in _D. orthogonia_ largely projecting.
Outer maxillæ, with their inner edges continuously covered with
bristles.
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