A Monograph on the Sub-class Cirripedia (Volume 1 of 2): The Lepadidae; Or, Pedunculated CirripedesDarwin, Charles
Science
A Monograph on the Sub-class Cirripedia (Volume 1 of 2): The Lepadidae; Or, Pedunculated Cirripedes
Darwin, Charles
Cirripedia; Lepadidae
_General Remarks._--As there are only two species as yet known, and as
these resemble each other in every respect most closely, a generic
description would be a useless repetition of the full details given
under _Ibla Cumingii_. I have taken this latter species as the type,
from having, owing to the kindness of Mr. Cuming, better and more
numerous specimens. Ibla and Lithotrya are the only two recent genera in
which the body of the animal is lodged within the peduncle; but there is
no distinction of any importance, though useful for classification,
between the capitulum and peduncle; and these two parts, as we have
seen, tend to blend together in some species of Conchoderma and Alepas.
The entire absence of calcareous matter in the valves and spines of the
peduncle, at first appears very remarkable; but we have seen a similar
fact in Alepas, and there is an approach to it in some varieties of
_Conchoderma aurita_ and _C. virgata_. In all four valves of Ibla, the
umbones, or centres of growth, are at their upper points. The horny
spines on the peduncle, are the analogues of the calcareous scales in
Scalpellum and Pollicipes; and in this latter genus, two of the species
have their scales, almost cylindrical, placed irregularly, with new ones
forming over all parts of the surface, and not exclusively at the
summit,--in which several respects there is an agreement with Ibla. The
shape of the body (_i. e._ thorax and prosoma, Pl. IV, fig. 8 _a´_) is
peculiar; but it is only a slight exaggeration of what we have seen in
several genera, and shall meet again in some species of Scalpellum. The
presence of hairs on the outer membrane of the prosoma is a peculiarity
confined to this genus amongst the Lepadidæ, though observed in the
sessile genus, _Chthamalus_. The caudal appendages in the _I.
quadrivalvis_ attain a greater length than in any other species of the
family, being four times the length of the pedicels of the sixth cirrus.
A far more important peculiarity is the fact of the oesophagus, in both
species, running over or exteriorly to the adductor scutorum muscle,
instead of, as in every other species, close under this muscle. I took
great pains in ascertaining the truth of this singular anomaly: the
course of the oesophagus is approximately represented in Pl. IV, fig. 8
_a´_ by faint dotted lines. The stomach has no cæca; the biliary folds
are longitudinal; there is a marked constriction at the line
corresponding with the junction of the thorax and prosoma. There are no
filamentary appendages.
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