A Monograph on the Sub-class Cirripedia (Volume 1 of 2): The Lepadidae; Or, Pedunculated CirripedesDarwin, Charles
Science
A Monograph on the Sub-class Cirripedia (Volume 1 of 2): The Lepadidae; Or, Pedunculated Cirripedes
Darwin, Charles
Cirripedia; Lepadidae
_Scuta_ slightly convex, thrice as long as broad; upper part much
acuminated; occludent margin almost straight; basal margin nearly at
right angles to the occludent margin; the tergal margin is separated
from the lateral margin by an angle more or less prominent; a slight
curved ridge runs from the umbo to this angle, and this deserves
especial notice, inasmuch as it indicates the outline which the valve
assumed in its earliest growth, and which is permanently retained in
most of the older fossil species. Along the occludent margin, there is a
trace of a ledge, developed in a variable degree, and which is noticed
only on account of the plainly visible ledge along this same margin, in
the allied genus Oxynaspis. The umbo, or centre of calcification, is
seated close to the occludent margin, and at about one fourth of the
length of the valve from the apex. Internally, (fig. 15, _a´_, Pl. V,)
the part above the umbo is flat; and beneath this upper part, there is a
large rounded hollow (_d_) for the adductor muscle: a fold or
indentation (_a_) running downwards from the umbo, extends in a very
oblique line across the occludent margin. This fold is of high interest
as giving lodgment to the Complemental Males, and will hereafter often
be referred to.
_Terga_, triangular, flat; occludent margin, very slightly arched.
_Carina_ much bent, with the umbo placed at barely one third of the
entire length of the valve from the apex. Two very slight ridges can be
perceived, one on each side, running from the umbo to the basal margin,
and separating the roof from the parietes of the valve; these ridges are
of great use in distinguishing the fossil carinæ of Scalpellum, from the
carinæ of Pollicipes. The part above the umbo is formed by the upward
production of a marginal slip along each side of the valve, which slips
in the fossil species (C in the woodcut, fig. 1, given in the
Introduction,) I have designated as the intra-parietes. The lower part
of the valve gradually widens from the umbo downwards; internally, the
whole is deeply concave, and continuously curved. The angle varies at
which the upper and lower portions externally meet each other; but is
never less than 135°. The upper part of the carina runs up between the
terga for three-quarters of their length; the basal margin does not
extend down low enough to pass between the carinal latera.
_Rostrum_, (fig. 15 _b´_, seen externally, and highly magnified,)
minute, almost hidden by the enveloping membrane and by the small
prominent umbones of the rostral latera; in area equalling about one
fourth of the rostral latera; externally pyramidal, with the upper side
rather longer than the lower; internally slightly concave, square, with
the upper margin and sometimes with the lower margin, slightly hollowed
out. Umbo of growth nearly central.
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