A Monograph on the Sub-class Cirripedia (Volume 1 of 2): The Lepadidae; Or, Pedunculated CirripedesDarwin, Charles
Science
A Monograph on the Sub-class Cirripedia (Volume 1 of 2): The Lepadidae; Or, Pedunculated Cirripedes
Darwin, Charles
Cirripedia; Lepadidae
The _Peduncle_ is of considerable length in some of the species, and
rather short in others; it is, in every case, clothed with calcified
scales. The scales in the first four species are placed alternately and
symmetrically; they are formed and added to in the same manner as in
Scalpellum; they differ in size according to the size of the individual,
and consequently the lower scales on the peduncle, formed when the
specimen was young, are smaller than the upper scales; the lower scales
are separated from each other by wide interspaces of membrane, owing to
the continued growth of the peduncle by the formation of new layers of
membrane, and the disintegration of the old outer layers. Each scale is
invested by tough membrane (or has been, for it is often abraded off),
in the same manner as the valves; each is furnished with one or more
tubuli, in connection with the underlying corium. In _P. sertus_ and _P.
spinosus_, the scales are small, spindle-shaped, and not of equal sizes,
and the rows are distant from each other, so that their alternate
arrangement is not distinguishable; in these two species, new scales are
formed round the summit of the peduncle, and the growth of each is
completed whilst remaining in the uppermost row; but, besides these
normal scales, such as exist in the other species of Pollicipes and in
Scalpellum, new scales are formed in the lower part of the peduncle,
which are generally of very irregular shapes, are often larger than the
upper ones, are crowded together, and sometimes do not reach the outer
surface of the membrane. This formation of scales in the lower part of
the peduncle, independently of the regular rows round the uppermost
part, is perhaps a feeble representation of the calcareous cup at the
bottom of the peduncle in the genus Lithotrya. The prehensile antennæ
will be described under _P. cornucopia_.
_Size._--Most of the species are large: and _P. mitella_ is the most
massive of the Pedunculated Cirripedes.
The _Mouth_ is not placed far from the adductor muscle. The labrum is
highly bullate. The mandibles have either three or four main teeth (Pl.
X, fig. 1), with often either one or two smaller teeth inserted between
the first and second. The maxillæ (Pl. X, figs. 13, 14), have their
edges either straight and square, or notched, or more commonly with two
or three prominences bearing tufts of finer spines. The outer maxillæ
(fig. 17) generally have a deep notch on their inner edges, but this is
not invariable. The olfactory orifices in most of the species are highly
prominent.
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