A Monograph on the Sub-class Cirripedia (Volume 1 of 2): The Lepadidae; Or, Pedunculated CirripedesDarwin, Charles
Science
A Monograph on the Sub-class Cirripedia (Volume 1 of 2): The Lepadidae; Or, Pedunculated Cirripedes
Darwin, Charles
Cirripedia; Lepadidae
The scutum overlaps either about one third or even one half of the
entire width of the tergum, and abuts against a prominent longitudinal
ridge on its exterior surface. In _L. truncata_ and _L. Valentiana_,
this ridge on the tergum being folded over towards the scutum, forms a
conspicuous furrow, receiving the tergal margin of the latter. In _L.
Valentiana_, there is a second furrow on the carinal side of the tergum,
receiving the upper end of the corium-covered or growing surface of the
carina. Besides these provisions for holding together the valves, there
are, apparently, others for a similar purpose; thus in each scutum,
under the rostral angle, there is a roughened knob-like tooth, which
touches the under side of the little rostrum, and no doubt serves to
give attachment to the membrane uniting the three valves together. In
some species, the adjoining basal margins of the scuta and terga, where
touching each other, are inflected and roughened; again in _L.
Rhodiopus_, the carinal angles of the terga are produced into points,
and in _L. truncata_ and _L. Valentiana_ into prominent roughened knobs,
which touch two corresponding small knobs, on the upper part of the
growing surface of the carina. Moreover, considerable portions of the
inner surfaces of the scuta and terga, are roughened with minute sharp,
imbricated points, apparently for the firmer attachment of the corium.
The roughened knobs at the rostral angles of the scuta, no doubt are
homologous with the teeth in a similar position on one or both scuta in
Lepas, and in some fossil species of Pollicipes, as in _P. validus_. The
other projections and roughened surfaces are peculiar to Lithotrya. The
growth of all the valves is, as in Pollicipes, simply downwards.
The _Scuta_ are triangular, with their umbones or centres of growth at
the apex; the tergal margin, as seen from within, is either nearly
straight or much hollowed out, accordingly as the scuta simply overlap
the terga, or are received in a furrow. In some of the species there is
a distinct pit for the adductor muscle, and in others this cannot be
distinguished.
_Terga._--These present great differences in shape; but all appear to be
modifications, (as seen internally,) of a rhomboidal figure, which seems
to be the normal form of the terga in the Lepadidæ. Of the lower part of
the valve, the whole exterior surface, with the exception of a narrow
ridge running from the apex down to the basal angle, is hidden by the
overlapping of the scuta, latera, and carina.
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