A Monograph on the Sub-class Cirripedia (Volume 1 of 2): The Lepadidae; Or, Pedunculated CirripedesDarwin, Charles
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A Monograph on the Sub-class Cirripedia (Volume 1 of 2): The Lepadidae; Or, Pedunculated Cirripedes
Darwin, Charles
Cirripedia; Lepadidae
I may here remark that, in most
sessile Cirripedes, the outside membrane connecting the operculum and
shell, is regularly moulted.
[20] The external integuments being moulted in Crustacea, but not
in the Cirripedia, may appear, at first, an important difference:
but we here see that non-exuviation is not universal amongst the
Lepadidæ, and, on the other hand, according to M. Joly, (‘Annales
des Sciences Naturelles,’ 2d series, Zoolog.), there is one true
crustacean, the _Isaura cycladoides_, which has a persistent
bivalve shell.
The delicate tunic lining the sack, (a mere duplicature of that thick
one, forming the outside of the capitulum, and generally transformed
into valves,) and the integuments of the whole body, are regularly
moulted. With these integuments, the membrane lining the oesophagus, the
rectum, and the deep olfactory pouches, and the horny apodemes of the
maxillæ, are all cast together. I have seen a specimen of Lepas, in
which, from some morbid adhesion, the old membrane lining one of the
olfactory pouches had not been moulted, but remained projecting from the
orifice as a brown shrivelled scroll. The new spines on the cirri (and
on the maxillæ) are formed within the old ones; but as they have to be a
little longer than the latter, and as they cannot enter these up to
their very points, their basal portions are not thus included, but are
formed, running obliquely across the segments of the cirri; and what is
curious, these same basal portions are turned inside out, like the
fingers of a glove when hastily drawn off. After the exuviation of the
old spines, the new spines have their inverted basal portions drawn out
from within the segments, and turned outside in, so as to assume their
proper positions.
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