A Monograph on the Sub-class Cirripedia (Volume 1 of 2): The Lepadidae; Or, Pedunculated CirripedesDarwin, Charles
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A Monograph on the Sub-class Cirripedia (Volume 1 of 2): The Lepadidae; Or, Pedunculated Cirripedes
Darwin, Charles
Cirripedia; Lepadidae
_Description._--Capitulum flattened, sub-triangular, composed of five
approximate valves. The valves are either moderately thick and
translucent, or very thin and transparent; and hence, though themselves
colourless, they are often coloured by the underlying corium. Their
surfaces are either smooth and polished, or striated, or furrowed, and
sometimes pectinated. They are not subject to disintegration; they are
generally naked, except on the borders, where they are coated, and held
together by membrane; in _L. fascicularis_, however, the valves are
covered with thin membrane, bearing very minute spines. The manner of
growth of the valves will be best described under each. All the valves,
even in the same species, are subject to considerable variation in
shape, more especially the terga.
_Scuta._--These valves are sub-triangular in outline, with the basal
margin straight and rather short; and with occludent and tergo-carinal
margins more or less protuberant; in _L. fascicularis_, however, the
basal (Pl. I, fig. 6), and occludent margins are slightly reflexed and
prominent. A ridge, generally runs from the umbo to the upper point.
Internally, there is no conspicuous pit for the adductor muscle; under
the umbones, there is generally either on both valves, or only on the
right-hand side (Pl. I, fig. 1 _c_), a small calcareous projection or
tooth, of variable size and shape, even in the same species; it is
generally largest on the right-hand valve; these teeth at first sight
appear to form a hinge, uniting the opposite scuta at their umbones, but
this is not really the case, and their use appears to be only to give
attachment to the membrane uniting the valves together, and to the
peduncle. The basal margin is internally strengthened by a calcified
rim, more or less developed. The umbones (and primordial valves when
distinguishable,) are seated at the rostral angles; during growth the
basal margin is not added to, and the occludent margin only to small
extent; hence the main growth of the valve is at the upper end, and
along the carina-tergal margin. In _L. fascicularis_, however, the basal
reflexed margin is slightly added to beneath the umbo.
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