A Monograph on the Sub-class Cirripedia (Volume 1 of 2): The Lepadidae; Or, Pedunculated CirripedesDarwin, Charles
Science
A Monograph on the Sub-class Cirripedia (Volume 1 of 2): The Lepadidae; Or, Pedunculated Cirripedes
Darwin, Charles
Cirripedia; Lepadidae
_Peduncle and Attachment._--The peduncle is generally quite smooth:
though with a high power its surface may be seen to be studded with
minute beads, or larger discs, of yellowish and hard chitine; in the
young of _L. australis_, and I suspect of some other species, it is
covered with very minute spines. The peduncle in this genus attains its
greatest development. The cement-tissue debouches, I believe, only
through the functionless larval antennæ, except in one species, _L.
fascicularis_, in which a ball of this substance is formed in a most
peculiar manner round the peduncle (Pl. I, fig. 6), apparently for the
purpose of serving as a float, as will be presently described.
_Size and Colour._--The species of this genus are the largest of the
Pedunculata, with the exception of some Pollicipes: even in the smallest
species (_L. pectinata_), the capitulum sometimes attains a length of
about half an inch. The peduncle varies much in length in the same
species: in _L. anatifera_, it is occasionally above a foot long. The
colours of _L. anatifera_, _L. Hillii_, and _L. anserifera_, are very
bright and striking; the membrane bordering the valves and that round
the top of peduncle in two of the species, is of the brightest
scarlet-orange; the valves, owing to the underlying corium, are pale
blueish-grey, and the interspaces between them dark leaden-purple. The
cirri and trophi are generally dark purple or lead-colour.
_Filamentary Appendages._--These are attached to beneath the basal
articulation of first pair of cirri; they vary in the several species,
from one to five or six on each side, the lowest being always the
longest. Several of them are occupied by testes. In _L. pectinata_,
generally, not even one is developed. They are subject to great
variation in their proportional lengths, and in number, in the same
species. These organs have generally been considered to serve as
branchiæ; I see no reason to believe that they are more especially
designed for this end, than is the general surface of the body.
_Mouth._--The labrum is moderately bullate, the longitudinal diameter of
this part equalling about one third, or half of that of the rest of the
mouth. The palpi are moderately developed. The mandibles (Pl. X, fig. 5)
have five teeth with the inferior point either broad, or very narrow and
tooth-like. The maxillæ are step-formed (Pl. X, fig. 9); the first step
is sometimes indistinct and curved; and in _L. pectinata_, all the steps
vary much, and are more or less blended together. The outer maxillæ
(like those at Pl. X, fig. 16), are internally clothed continuously with
spines. The olfactory orifices are not at all prominent.
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