A Monograph on the Sub-class Cirripedia (Volume 1 of 2): The Lepadidae; Or, Pedunculated CirripedesDarwin, Charles
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A Monograph on the Sub-class Cirripedia (Volume 1 of 2): The Lepadidae; Or, Pedunculated Cirripedes
Darwin, Charles
Cirripedia; Lepadidae
_Size._--All the species are small, with a capitulum not exceeding half
an inch in length.
_Filamentary Appendages._--None.
_Mouth._--Labrum generally considerably bullate in the upper part, with
a row of teeth on the crest. The _mandibles_ have four teeth, with the
inferior point narrow and spine-like, or rudimentary and absent. The
_maxillæ_ have, under the two or three upper great spines, a deep notch
itself bearing spines; beneath this, the lower part is straight and
considerably prominent, Pl. X, fig. 15. Outer maxillæ are covered on
their inner sides continuously with spines.
_Cirri._--The first pair is sometimes seated very distant from the
second. The arrangement of the spines on the posterior cirri varies, to
an unusual degree within the limits of the same genus. We have either
the ordinary structure of anterior pairs, with single fine intermediate
spines (as in _P. Kæmpferi_ and _aurantia_), or we have the pairs
increased by one or two additional longitudinal lateral rows, as in _P.
eburnea_; or we have the front spines forming a single transverse row,
as in _P. crassa_ and _P. fissa_, Pl. X, fig. 29, _a_. The segments in
none of the species are protuberant; the anterior ramus of the second
cirrus does not seem to be thicker than the posterior ramus, as is
usually the case. The rami of the second, and of most of the other
cirri, are unequal in length,--the anterior ramus, contrary to the
ordinary rule, being longer in _P. eburnea_, _P. fissa_, and _P.
crassa_, than the posterior ramus by several segments; I have hitherto
observed this inequality only in the sessile genus Chthamalus.
The _Caudal Appendages_ are small, uniarticulate, and always furnished
with bristles.
_Distribution._--Four out of the five species live attached to
Crustacea in the European and Eastern warmer temperate and
tropical oceans; the fifth species was found attached to the
dead spines of an Echinus, off New Guinea. It is probable that
several more species will be hereafter discovered.
1. PÆCILASMA KÆMPFERI. Pl. II, Fig. 1.
_P. valvis 5; carinæ basi truncatâ et cristatâ: scutorum dentibus
internis umbonalibus fortibus: tergorum acumine basali truncato,
margini occludenti pæne parallelo._
Valves 5; carina with a truncated and crested base; scuta with strong
internal umbonal teeth; terga with the basal point truncated, almost
parallel to the occludent margin.
Maxillæ with short thick spines in the notch under the two upper great
spines; caudal appendages with scattered bristles on their summits, and
along their whole outer margins.
Japan; attached, in great numbers, to the upper and under sides
of the _Inachus Kæmpferi_ of De Haan, a slow-moving brachyourous
crab, probably from deep water. British Museum.
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