A Monograph on the Sub-class Cirripedia (Volume 2 of 2): The Balanidæ, (or Sessile Cirripedes); the Verrucidæ, etc., etc.Darwin, Charles
Science
A Monograph on the Sub-class Cirripedia (Volume 2 of 2): The Balanidæ, (or Sessile Cirripedes); the Verrucidæ, etc., etc.
Darwin, Charles
Balanidae; Cirripedia
The drawing in Pl. 30, fig. 8, of part of the second segment, of the
third or disc segment, and of the fourth or ultimate segment, in
_Lepas australis_, is, I think, very accurate. The second segment
articulates on the upper or dorsal surface of the disc, and has the
articulation on one side constricted and formed of thin and flexible
membrane; the little terminal segment, which is turned outwards at
right angles, also, articulates on the disc. That the disc forms a
true segment is shown clearly in Cryptophialus (Pl. 24, fig. 18),
where the articulation with the second segment is not in such close
contact. The disc is either circular, as in Lepas, or hoof-shaped, as
in Ibla: in _B. balanoides_ the disc is rather hollowed out on the
inner side. It has the power of adhering even to so smooth a substance
as glass, placed vertically. It is surrounded by a rim of transparent
membrane. On the hinder margin some spines arise from the central and
more opaque part: in _Lepas australis_, there are no less than seven of
these spines (fig. 8): in _Conchoderma virgata_ there are only four,
in _Scalpellum_ and _Ibla_ only one. When the disc is placed on the
surface of attachment, these spines lie parallel to it. The middle
part of the disc is, almost always, nearly opaque; and in tracing
the cement-ducts from within the body of the pupa, or of the young
Cirripede, I in many cases traced them as far as this point, but here
lost them. From this same obscure central part of the disc, in most,
if not in all species of the Lepadidæ, spokes radiate, which sometimes
are branched, and are not regular, not always even resembling each
other on the opposite sides of the same individual. Round the proper
membranous border of the disc, a second one may be observed (fig. 8,
_p_), which differs in shape and extent in different specimens: under
favorable circumstances, and very high powers, it may be seen to
have a reticulated structure, and to be of a very pale brown colour;
towards the exterior margin, the reticulations become finer, and are
blended together and lost; on the inner margin, the substance forming
this membrane may be seen to come out of the spokes. This substance
is the cement, which has the power of adhering to whatever substance
it grows against; and thus the disc of each antenna becomes cemented
down, and soon both the antennæ are surrounded by a common border of
cement, which gradually increases, after the metamorphosis, in extent.
Occasionally the cement forms little projections, like short spines, on
the edges opposite to the orifices of the spokes.
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