A Monograph on the Sub-class Cirripedia (Volume 2 of 2): The Balanidæ, (or Sessile Cirripedes); the Verrucidæ, etc., etc.Darwin, Charles
Science
A Monograph on the Sub-class Cirripedia (Volume 2 of 2): The Balanidæ, (or Sessile Cirripedes); the Verrucidæ, etc., etc.
Darwin, Charles
Balanidae; Cirripedia
On the _internal viscera_ I have nothing to add. The cement-duct is
represented in Pl. 30, fig. 2, _t′_, on the near side, running into
the antennæ; and I repeatedly traced it, for the duct is very strong,
as far as the disc segment; at the other end it joins the cement-gland
(_t_) on the same side of the body. This cement-gland is proved, by
the clearest series of facts, to be converted into the incipient
ovaria and ovarian cæca. The cement-glands in the older pupæ could be
traced as far as the cæca of the stomach, exactly where the ovaria lie
in the mature animal; but in some young pupæ, they extended further
posteriorly, past the mouth, between the outer and inner membranes
of the overlapping carapace. I have faintly shown the course of the
stomach, with its two cæca at the upper end; the anus lies between the
caudal appendages, at the extremity (above _b′_) of the abdomen. At
this age there is no trace of the vesiculæ seminales, so conspicuous in
the mature Cirripede.
_Young Cirripede, whilst within the pupa._--I repeatedly succeeded in
dissecting the young _Lepas australis_ out of the pupa; and by the
previous action of boiling potash, and by a strong light, I was enabled
to make a camera sketch (Pl. 30, fig. 2) of the relative positions of
their several parts. The young Cirripede is drawn very faintly, and is
best seen by holding the plate in the same position with the mature
animal, of which a section is given in my volume on the Lepadidæ, Pl.
9, fig. 4. I may just notice how complicated are the membranes in a
longitudinal section taken at this period: we have, 1st, beginning at
the back, the horny tissue of the carapace or bivalve shell of the
pupa; 2d, the primordial valve (_z_, in fig. 3) of the young Cirripede;
3d and 4th, two folds of corium; 5th, the membrane of the sack of the
Cirripede; 6th, the membrane of the sack of the pupa; 7th, the outer
tunic of the thorax of the pupa; 8th, the outer tunic of the thorax
of the young Cirripede; 9th, the corium lining the latter membrane;
and these nine membranes would be repeated on the opposite side of the
section, if it were taken through either side of the shell or carapace,
bordering the orifice.
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