A Monograph on the Sub-class Cirripedia (Volume 2 of 2): The Balanidæ, (or Sessile Cirripedes); the Verrucidæ, etc., etc.Darwin, Charles
Science
A Monograph on the Sub-class Cirripedia (Volume 2 of 2): The Balanidæ, (or Sessile Cirripedes); the Verrucidæ, etc., etc.
Darwin, Charles
Balanidae; Cirripedia
_Compartments._--The external appearance of the shell has already been
described. In the most typical species, the parietes consist of an
outer and inner lamina, separated by strong longitudinal septa; these
septa are denticulated on both sides at their bases, but only close
to the inner lamina; in fact the inner lamina is apparently formed by
the union, thickening, and production, of some of the denticuli. As
it is not the innermost of the denticuli on the basal edges of the
longitudinal septa, which thus become united into a solid layer, the
longitudinal septa form slightly projecting, longitudinal ribs on the
inner lamina. These internal ribs are longitudinally striated; in
old specimens they often become obliterated, especially in the upper
part of the shell. The parietal tubes or pores (occupied by threads
of corium) are generally square and large; but in _B. Ajax_ they
are very small, and in _B. glandula_ often extremely minute. In the
upper part of the shell, and sometimes low down, they are generally
crossed by thin, transverse, calcareous septa: in some species, as
in _B. perforatus_, and in some varieties of _B. amphitrite_, the
upper ends of the tubes are filled up solidly with shell. In some
varieties of _B. crenatus_ and of _amphitrite_, the longitudinal septa,
near the outer lamina, divide, thus giving rise to a very imperfect
row of outer short tubes. In _B. vinaceus_ (Pl. 2, fig. 7 _d_) the
inner lamina is cancellated instead of being solid, which is caused
by the basal denticuli of the longitudinal septa being simply united
together by their ends and crossed by transverse septa, instead of
being consolidated into a mass. In several species, as in _B. Hameri_,
the walls consist only of the outer lamina with longitudinal ribs, no
inner lamina having been formed; the ribs here evidently answer to the
longitudinal septa in the foregoing species. In _B. flosculus_ and
_imperator_ the walls are solid, their basal margins being formed of
irregular, elongated points, and little ridges (Pl. 8, fig. 4 _c_),
which apparently prefigure the more regular longitudinal ribs or septa.
In _B. balanoides_ the walls are generally either nearly smooth and
solid, or irregularly cancellated; in _B. cariosus_ (Pl. 7, fig. 3 _b_)
two or three rows of short irregular tubes are formed by unequally
branching septa, almost as in the genus _Tetraclita_.
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