A Monograph on the Sub-class Cirripedia (Volume 2 of 2): The Balanidæ, (or Sessile Cirripedes); the Verrucidæ, etc., etc.Darwin, Charles
Science
A Monograph on the Sub-class Cirripedia (Volume 2 of 2): The Balanidæ, (or Sessile Cirripedes); the Verrucidæ, etc., etc.
Darwin, Charles
Balanidae; Cirripedia
express my deliberate opinion, that every part and organ, internal and
external, in Cirripedes, is liable to some amount of variation in some
of the species.
I must now point out the principal changes which supervene during
growth, and which cannot properly be called variations. In the first
place, I think, it is scarcely possible to recognise a species when
under the 1/10th of an inch in diameter. In some cases, as in _var.
d'Orbignii_ of _B. tintinnabulum_, the shell is invariably coloured
when old, but quite white when very young. Generally the tints become
very much darker with age. Some species, which usually or invariably
have, when mature, longitudinally folded walls, as _B. flosculus_
and _balanoides_, are perfectly smooth in youth. The walls in _B.
eburneus_, when young, have white, hyaline, longitudinal lines, and
are naked; whereas, with advancing age, these lines disappear, and the
subsequently formed shell becomes covered with membrane. The summits
of the radii are apt to be oblique in the young of _B. Capensis_,
_psittacus_, and _tintinnabulum_, whereas they are generally quite
square in old specimens. In _B. eburneus_, _cariosus_, and in a lesser
degree in _B. psittacus_, the scuta become longitudinally striated
only with age. On the other hand, in very young specimens of _B.
tintinnabulum_, the scuta sometimes are deeply impressed by little
pits placed in rows. I have already alluded to the longitudinal
furrow on the tergum so entirely changing its character, owing to the
edges becoming, during growth, folded inwards; this likewise causes a
decrease in the proportional breadth of the spur. In old specimens of
_B. flosculus, var. sordidus_, the whole tergum is much more elongated
than in young specimens. The basal margin of the sheath is hollow
beneath in the young of _B. cariosus_ and of some other species, but
in the old it is continuous with the inner surface of the walls. The
inner lamina of the parietes generally loses, to a certain extent, its
longitudinally ribbed character in old age. The basis is solid, instead
of being porose, in very young specimens of _B. improvisus_. In all the
species, the carino-lateral compartments, in early age, are very narrow
in proportion to the width of the lateral compartments; and in all, at
this early period, the operculum is large in proportion to the whole
shell. The number of spines on the edge of the maxillæ, the number of
segments in all the cirri, and the number of spines on each segment,
are few in early youth, and go on increasing with each successive
exuviation: the pedicels of the cirri are long in proportion to the
rami at this same early age.[84]
[84] In some specimens of _Balanus perforatus_ I made the following
enumeration of the number of segments in the cirri:--
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