A Monograph on the Sub-class Cirripedia (Volume 2 of 2): The Balanidæ, (or Sessile Cirripedes); the Verrucidæ, etc., etc.Darwin, Charles
Science
A Monograph on the Sub-class Cirripedia (Volume 2 of 2): The Balanidæ, (or Sessile Cirripedes); the Verrucidæ, etc., etc.
Darwin, Charles
Balanidae; Cirripedia
_Fossil Species._--Having already given, under the Family, some account
of the geological history of sessile cirripedes, short as it is, I here
only allude to the subject in order to state my conviction that species
cannot be satisfactorily distinguished in a fossil state, and rarely
in a recent state, without an examination of the opercular valves.
Nothing, indeed, could have been easier than to have affixed names
to many groups of specimens, having different aspects, but to feel
sure that these were really distinct species requires better evidence
than can be afforded by the shell, without the operculum. No doubt,
in some of the smaller sections of the genus--for instance, in that
characterised by a membranous basis--it would have been possible to
have distinguished some or several fossil species; but such have not as
yet been found. When the specimens are much fossilised, it is, indeed,
difficult to make out the primary points of structure--namely, whether
the parietes, radii, and basis are porose: to do this it is sometimes
necessary to rub down, polish, and carefully examine, a transverse
section of a piece of the shell.
_Sections of the Genus._
[A]
_Parietes, and basis, and radii permeated by pores._
[B]
_Parietes and basis sometimes permeated by pores, sometimes not; radii
not permeated by pores; shell elongated in its rostro-carinal axis;
basis boat-shaped, attached to Gorgoniæ and Milleporæ._
[C]
_Parietes and basis permeated by pores; radii not permeated by pores._
[D]
_Parietes permeated by pores. Basis and radii not permeated by pores._
[E]
_Basis membranous._
[F]
_Parietes and radii not permeated by pores. Basis sometimes permeated
by pores, sometimes not, sometimes excessively thin and hardly
distinguishable._[86]
[86] The following species might, owing to variation, or to the
obscurity of their structure, without care, be classed in the wrong
sections:--in A, _Balanus Ajax_, from living attached to Milleporæ,
is sometimes elongated in its rostro-carinal axis, and from having
its radii only finely porose, it might be classed in B: on the
other hand, _Bal. stultus_ is sometimes but little elongated, and
the basis hardly boat-formed, and hence might be classed in A. The
distinction between all the species in (B) and the sub-genus Acasta
is artificial.
In sections C and D, I have seen one specimen of _B. spongicola_
with a solid basis, and very young specimens of _B. improvisus_
are thus characterised, and therefore these species are liable to
be placed in the wrong section, D: _Bal. nubilus_, also, has part
of its base non-porose, and therefore likewise might be placed
in D: on the other hand, the circumference of the basis in _B.
patellaris_ is often porose, and hence this species, which belongs
to D, might be placed in C.
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