A Monograph on the Sub-class Cirripedia (Volume 2 of 2): The Balanidæ, (or Sessile Cirripedes); the Verrucidæ, etc., etc.Darwin, Charles
Science
A Monograph on the Sub-class Cirripedia (Volume 2 of 2): The Balanidæ, (or Sessile Cirripedes); the Verrucidæ, etc., etc.
Darwin, Charles
Balanidae; Cirripedia
comes to be thus invested; and is marked by circular lines, one above
the other, caused by the successive exuviations. This investing
membrane often supports rows of minute bristles, directed upwards.
Generally, a film of shell is deposited, at the period of the formation
of each new opercular membrane, on that part of the sheath which lies
immediately beneath. This innermost film or thin layer of shell, on
the lines of suture between the compartments, breaks joint, at least
in some cases, with the underlying shelly layers,--that is, the suture
in this last-formed film does not lie exactly over the suture in the
subjacent layers of the sheath. In Tubicinella, the sheath extends down
close to the basis; and what is unique in this one genus, the opercular
membrane, gradually thinning out downwards, closely adheres to the
whole inner surface of the shell. In Tubicinella and in Xenobalanus
(Pl. 17, fig. 4 _b_), the sheath separates easily into separate
successive rims of shell; and this structure evidently is for the sake
of facilitating the breakage of the upper end of the shell, which, as
we shall presently see, is necessary to allow of the increase in size
of its orifice.
_Structure of the Basis._
This, in several genera and species, is composed of simple membrane,
and consists of successive, concentric, circular slips, added round the
outside, at each period of growth. In some species of Tetraclita and
Balanus the basis is calcareous, but diaphanous, very thin, smooth, or
somewhat granulated. In other cases it consists of a single calcareous
lamina, either smooth, or with ridges radiating from its centre; it
is formed of two laminæ, (as is most usual in Balanus,) separated by
radiating septa. These septa, as well as the radiating ridges in the
case of the single lamina, are homologous with the longitudinal septa
of the parietes. The denticulated ends of the latter enter the mouths
of the tubes formed by the radiating septa of the basis: threads of
corium pass between the denticuli of the parietal septa, and thus
enter the basal tubes. The ends of these threads of corium generally
deposit transverse calcareous septa, exactly as within the parietal
tubes. When the basis is thick the septa themselves (_ccc_) between
the proper basal tubes, become porose, (or rather cancellated,) and
they sometimes expand into a very thick, cancellated layer, separating
the outer lamina (_a_) of the basis from the proper basal tubes, which
always lie close under the inner lamina (_b_). This structure differs
only slightly from that seen in the parietes of Tetraclita, in which
the branching of the longitudinal parietal septa, produces thick walls,
formed of several rows of tubes or pores. With respect to peculiarities
in structure of the basis, _Balanus lævis_ offers the most remarkable
case; for here, in specimens which have grown crowded together, the
whole interior appears sometimes to have become too much elongated or
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