A Monograph on the Sub-class Cirripedia (Volume 2 of 2): The Balanidæ, (or Sessile Cirripedes); the Verrucidæ, etc., etc.Darwin, Charles
Science
A Monograph on the Sub-class Cirripedia (Volume 2 of 2): The Balanidæ, (or Sessile Cirripedes); the Verrucidæ, etc., etc.
Darwin, Charles
Balanidae; Cirripedia
too deep for the animal's body, and consequently the lower part of
the deeply-concave basis has been filled up (Pl. 4, fig. 2 _a_) by
thin, irregular, calcareous diaphragms. In elongated specimens, also,
of _Balanus balanoides_, the shell sometimes appears to have grown
too long for the animal's body; but in this case the membranous basis
becomes extremely convex inwards; it still reaches the basal edges of
the parietes all round, but in the middle it is raised high above the
surface of attachment; yet sometimes threads of the cementing tissue
depend from the middle part to the surface of attachment. In _Balanus
terebratus_ (Pl. 8, fig. 2 _a_, 2 _b_), and in some species of Acasta,
the basis is riddled, as previously stated, by numerous, minute,
membrane-covered orifices. In _B. declivis_ the membranous basis is
always extremely oblique, owing to the rostral end of the shell being
twice as high as the carinal and opposite end.
[Illustration: Fig. 11.
Portion of edge of basis of _Balanus tintinnabulum_, _a_, _a_, outer
lamina; _b_, _b_, inner or upper lamina; _c_, _c_, _c_, porose or
cancellated radiating septa.]
Regarding the very remarkable means by which the basis of sessile
Cirripedes is cemented to the surface of attachment, it will be
convenient to defer for a little the description, on account of its
necessary length.
_Structure of the Opercular Valves (Scuta and Terga)._
These are situated on each side of the slit or orifice leading into the
sack; from their shape, their powers of movement, their separation by
flexible membrane from the shell, to which they serve as a lid, they
appear at first as if they constituted an element very distinct from
the shell itself, but this is not the case. They are, together with the
opercular membrane, as essentially as the whole of what is externally
visible, a part of the modified carapace, of which they occupy the
upper or posterior extremity: from tracing the metamorphoses, or
even by comparison of a Balanus with Pollicipes, there can be no
doubt of the truth of this conclusion. The opercular valves are four
in number,--a pair of scuta and a pair of terga; but the latter in
_Coronula diadema_ and _reginæ_, are either aborted or represented by
a mere rudiment; and in Xenobalanus both scuta and terga are quite
absent. In several cases, more especially in the genus Pyrgoma (Pl.
13, fig. 1 _b_), the scutum and tergum on each side are calcified
together, so that sometimes not even a trace of the line of junction
can be discovered. In most cases the scutum is firmly united, being
articulated in a manner presently to be described, to the tergum; but
in Coronula, Tubicinella (Pl. 17, fig. 3 _c_), and Platylepas, the ends
of these valves are simply approximated.
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