A Monograph on the Sub-class Cirripedia (Volume 2 of 2): The Balanidæ, (or Sessile Cirripedes); the Verrucidæ, etc., etc.Darwin, Charles
Science
A Monograph on the Sub-class Cirripedia (Volume 2 of 2): The Balanidæ, (or Sessile Cirripedes); the Verrucidæ, etc., etc.
Darwin, Charles
Balanidae; Cirripedia
The opercular valves are added to along their basal margins alone;[23]
the animal's body, together with the several muscles, becoming attached
at each period of growth lower and lower down to the valves; this
no doubt is effected by the absorption of the upper surfaces of the
muscles, and the formation of new fasciæ on their lower surfaces. The
opercular membrane, which, though thin and flexible, forms part of
the general outer surface of the animal as much as does any portion
of the rigid shell, with which indeed it is strictly homologous, is
periodically moulted, together with the integuments of the whole
included animal. The new opercular membrane is of course each time
formed a little larger than the old one. In Coronula and Tubicinella,
however, several successive opercular membranes are preserved one
over the other, and the outside membrane gradually disintegrates; in
these cases the undermost opercular membrane is formed wrinkled and
considerably too large, so as to allow of being stretched, before it
is finally cast off. In Tubicinella, the opercular membrane runs down,
adhering to the inner surface of the shell, to nearly the basis, and
hence during the diametric growth of the shell, it is longitudinally
split, and is repaired by slips of new membrane, which resemble the
radii in form and in direction of the lines of growth.
[23] In some species of Pyrgoma, the ledge (_limbus occludens_)
which is added along the occludent margin of both scuta and terga,
and in some species of Balanus a narrow rim, or slight protuberance
which is added along the carinal margin of the terga, offer
unimportant exceptions to the rule, that the opercular valves grow
only at their basal margins.
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