A Monograph on the Sub-class Cirripedia (Volume 2 of 2): The Balanidæ, (or Sessile Cirripedes); the Verrucidæ, etc., etc.Darwin, Charles
Science
A Monograph on the Sub-class Cirripedia (Volume 2 of 2): The Balanidæ, (or Sessile Cirripedes); the Verrucidæ, etc., etc.
Darwin, Charles
Balanidae; Cirripedia
In my former volume on the Lepadidæ, I have described the _ovigerous
fræna_ occurring on the two sides of the sack, to which the
_ovigerous lamellæ_ are attached by a peculiar glandular secretion:
in the Balanidæ there are no ovigerous fræna, but the branchiæ just
described are identical with the fræna in essential structure and
in position; differing only in being placed a little nearer to the
carinal end of the sack, and in being generally (but not always)
larger and more plicated: seeing this, and that in _Alcippe lampas_,
and in some species of Pollicipes,--the genus which comes nearest to
the Balanidæ,--the ovigerous fræna are large and are destitute of
glands, and have therefore lost their normal function of supporting
the ovigerous lamellæ, I can hardly doubt that the _branchiæ_ in the
Balanidæ are the _ovigerous fræna_ of the Lepadidæ in a modified
condition; a transformation of function not greater than that of the
swimming bladder of a fish into the lungs of the higher Vertebrata.[29]
[29] There is, I conceive, no foundation for the belief of some
authors that the branchiæ of the Balanidæ are in any way connected
with the ovaria.
_Thorax and Body._
_Parts of the body included within the shell or carapace._--These
parts (Pl. 25, fig. 1) consist of the prominent mouth, and of the
thorax (_c′_), with its largely developed portion, called the prosoma
(_c_), and with its appendages. The abdomen is quite rudimentary,
being represented merely by a small portion of membrane surrounding
the anus, and sometimes inserted like a wedge between the inwardly
inflected posterior thoracic segments; in only two genera (Catophragmus
and Pachylasma), its nature is rendered somewhat plainer by supporting
caudal appendages. The probosciformed penis lies folded under the
thorax; and I believe (from what is seen in the anomalous genus
Proteolepas), that it normally arises from the ventral surface of
the terminal point of the rudimentary abdomen.[30] The thorax is
laterally compressed, the ventral surface being very narrow, with
the bases of the cirri placed closely together. It consists, in
appearance, of two very different portions; one a soft, more or less
rounded bag, which I have called the prosoma; and the other, which
supports the five posterior pairs of cirri, is narrower, invested
with stiffer membrane, and is more or less distinctly composed of
five segments. These segments (Pl. 26, fig. 8) on their dorsal and
dorso-lateral surfaces, are generally driven like wedges one into the
other, with their points directed anteriorly: on the ventral surface
the articulations are transverse. The prolongation (_e_) of the thin
membrane (_a_) surrounding the anus (_b_), that is, the rudiment of
the abdomen, which sometimes carries caudal appendages, almost divides
(in appearance, whether really I know not) the hindermost thoracic
segment along the medio-dorsal line, into two parts. I have given the
Public-domain text, read in full here on John Shaqi.
Reviews
Reviews
No reviews yet
Be the first to share your thoughts on this work.
Elsewhere in the archive
Join the Discussion
Join the discussion
Sign in to leave a comment or review.
Sign InorCreate an account