A Monograph on the Sub-class Cirripedia (Volume 2 of 2): The Balanidæ, (or Sessile Cirripedes); the Verrucidæ, etc., etc.Darwin, Charles
Science
A Monograph on the Sub-class Cirripedia (Volume 2 of 2): The Balanidæ, (or Sessile Cirripedes); the Verrucidæ, etc., etc.
Darwin, Charles
Balanidae; Cirripedia
movement, are all admirably adapted to secure any prey, when once
entangled by the cirri.
_Mouth._
The mouth, in the sub-family Chthamalinæ, cannot be distinguished
from that of the Lepadidæ, which has been pretty fully described
in my former volume. In the Balaninæ, however, the labrum differs
considerably in not being swollen; that is, in its outer and inner
fold of membrane being close together, and in having a central notch:
the palpi are also larger, and the lower teeth on the mandibles, are
laterally (Pl. 26, fig. 5) double, as will be more particularly stated
under these two sub-families. I have given a drawing (Pl. 26, fig.
1) of the mouth, seen from above, of _Balanus perforatus_, with the
right-hand palpus (_d′_) and outer maxilla (_a′_) cut off, in order
that the labrum (_e_), mandibles (_c_), and inner maxilla (_b_) might
be better shown; the cut-off bases (_x_, _x_) of the first cirrus on
each side are also shown. In fig. 2 we have the deep supra-œsophageal
cavity in _Bal. improvisus_ torn open and laid flat, with the inner
surfaces of the labrum (_c_) and outer maxillæ (_a_) exhibited, the
palpi, mandibles, and inner maxillæ having been removed. Figs. 3 and
4 will presently be referred to; they are parts of the mouth, with
the muscles, &c. removed, of Coronula. The mouth differs extremely
little in the different genera and species of the Balanidæ, much less
than amongst the Lepadidæ. In the Balaninæ, the crest of the labrum
is sometimes hairy, instead of having, as is usual, from two to six
teeth on each side of the central notch: in _Balanus improvisus_ (Pl.
26, fig. 2) and _eburneus_, and in _Chelonobia_, the crest on each
side of the central notch (_e′_) is furnished with a row of finely
graduated teeth. A sub-triangular portion of the inner fold of membrane
of the labrum, which overhangs the œsophagus, is always thickened and
yellowish; it is also often punctured in patterns (Pl. 26, fig. 2,
_f_), which, I believe, give attachment to little muscles that serve
to open the upper end of the œsophagus. Opposite to this thickened,
sub-triangular portion of membrane, the thin membrane forming the
supra-œsophageal cavity (or the cavity surrounded by the gnathites)
is strengthened by a pair of curved ribs (_h_, fig. 2) of thickened
yellowish membrane, running down from the inner bases (_a′′_) of the
bilobed outer maxillæ to the opening of the œsophagus (_g_): a broad
branch from each of these ribs supports the sides of the orifice of
the œsophagus; and this branch almost joins on to a slightly thickened
rim or bar (_f′_), which branches off from the upper part of the
sub-triangular (_f_) inner fold of the labrum. This structure, in _Bal.
improvisus_, is represented in Pl. 26, fig. 2, as well as it could be,
considering that the deep supra-œsophageal cavity has to be torn open;
and then laid flat.
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