A Monograph on the Sub-class Cirripedia (Volume 2 of 2): The Balanidæ, (or Sessile Cirripedes); the Verrucidæ, etc., etc.Darwin, Charles
Science
A Monograph on the Sub-class Cirripedia (Volume 2 of 2): The Balanidæ, (or Sessile Cirripedes); the Verrucidæ, etc., etc.
Darwin, Charles
Balanidae; Cirripedia
the palpi: the point of union, when viewed from the outside, is seen
to form a knob on the shoulder of the labrum, beneath the level of
its crest, and at this knob (Pl. 26, fig. 3, close to _d′_) several
thickened bands in the surrounding membrane unite. The free portion of
the palpus stands out transversely behind (_i. e._ anteriorly to, in a
homological sense) the labrum. I suspect that the palpus possibly may
consist of two segments, of which the terminal one is free, and the
lower one confluent with the labrum.
Before proceeding any further, I should observe that figs. 3 and 4, in
Pl. 26, represent the membranes of the mouth of _Coronula diadema_,
perfectly cleaned. In fig. 3, all the front part of the mouth has been
removed, the mandible on one side, the labrum with the two palpi, and
the œsophagus being alone left, and these are viewed from the inner
side; the front part, however, of the supra-œsophageal cavity has been
cut away. In fig. 4, the labrum, with the œsophagus, has been removed,
whilst the two outer maxillæ, the right-hand inner maxilla and mandible
(with the exterior and basal portions, _d_, _d′′_, of one palpus) are
seen from the outside; but in order that these parts should all be
shown, the whole of the right-hand side of the mouth has been spread
out, for the teeth of the mandible should have stood in a vertical
line between the two outer maxillæ. In the mandibles, the free upper
part is separated, by a distinct articulation, from the square piece
of thickened membrane (fig. 3, _c^1_) on which it is supported; and
this latter is separated by a second articulation from a portion of
thickened membrane (_c^2_), the basal edge of which forms the third and
lowest articulation, separating the mouth from the body. This basal,
thickened portion of membrane curls round and inwards, towards the
outer maxillæ or front of the mouth, and its terminal points sometimes
even penetrate a little way within the muscles, like apodemes: it is
not distinctly separated by any line or suture from the membrane, which
forms the whole broad labrum; so that I at first concluded that the
labrum dipped under the mandibles, and thus afforded a support on which
they were articulated; but this appears so opposed to all analogy,
that it is more probable that the above basal thickened portion of
membrane is truly the basal segment of the mandibles, completely
confluent with the labrum; and it is, I think, not very improbable
that even a large portion of what in appearance belongs to the labrum,
namely, those concavities to which the muscles of the mandibles are
attached, may, also, be part of the basal segment of the mandibles.
Whether or no there really are two segments beneath the upper free
portion of the mandibles, which have become laterally confluent with
other parts, I must think that the square thickened piece of membrane
(_c^1_) represents at least one segment. I may here observe, that Prof.
Public-domain text, read in full here on John Shaqi.
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