A Text-book of Entomology: Including the Anatomy, Physiology, Embryology and Metamorphoses of Insects for Use in Agricultural and Technical Schools and Colleges as Well as by the Working EntomologistPackard, A. S. (Alpheus Spring)
Science
A Text-book of Entomology: Including the Anatomy, Physiology, Embryology and Metamorphoses of Insects for Use in Agricultural and Technical Schools and Colleges as Well as by the Working Entomologist
Packard, A. S. (Alpheus Spring)
Insects
The act of folding the broad hind wings among beetles is not less
significant than the arrangement of the fore wing. If we forcibly
spread out the former in a beetle which has just been killed and
then leave it to its own resources again, we observe the following
result: According to its peculiar mode of joining, the costal vein
on the fore edge approaches the mid or discoidal vein of the basal
half as well as the distal half of the wing, whence arises a
longitudinal fold which curves in underneath. Then the distal half
snaps under like the blade of a pocket knife and lies on the plane
of the costal edge of the wing, while it also draws after it the
neighboring wing-area. The soft hinder-edge portion turns in
simultaneously when this wing-area remains fixed to the body while
the costal portion is moving towards the middle line of the body.
The wing-membranes of almost all insects have, moreover, the
capability of folding themselves somewhat, and this power of
extending or contracting the wing-membrane at will is of great
importance in flight.
Yes, but how is the folded wing spread out again? The fact may be
shown more simply and easily than one might suppose, and may be most
plainly demonstrated even to a larger public by making an artificial
wing exactly after the pattern of the natural one, in which bits of
whalebone may take the place of veins and a piece of india rubber
the membrane spread out between them. The reader will be patient
while we just explain to him the act of unfolding of the membranous
wing of the beetle. The actual impulse for this unfolding is due to
the flexor muscles which pull on, and at the same time somewhat
raise the vein on the costal edge. By this means the membranous fold
lying directly behind the costal vein is first spread out. But since
this fold is connected with the longitudinal fold of the distal end
of the wing which closes like a blade, the wing-area last mentioned
which is attached to the middle fold of the wing by the elastic
spring-like diagonal vein becomes stretched out. The hinder rayed
portion adjacent to the body is, on the other hand, simply drawn
along when the wing stands off from the body.
In order to properly grasp the mechanism of the insect wing we must
again examine its mode of articulation to the body somewhat more
accurately.
[Illustration:
FIG. 171.—Longitudinal section through a Tipula: _a_, mouth; _an_,
antenna; _k_{3}_, maxillary palpus; _ol_, labrum; _oG_, brain;
_uG_, subœsophageal ganglion; _BG_, thoracic ganglion; _schl_,
œsophagus; _mD_, digestive canal; _Ov_, ovary; _vF_, fore wing;
_sch_, halter; _lm_, longitudinal—_b-r_, lateral muscles.—After
Graber.
]
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