A Text-book of Entomology: Including the Anatomy, Physiology, Embryology and Metamorphoses of Insects for Use in Agricultural and Technical Schools and Colleges as Well as by the Working EntomologistPackard, A. S. (Alpheus Spring)
Science
A Text-book of Entomology: Including the Anatomy, Physiology, Embryology and Metamorphoses of Insects for Use in Agricultural and Technical Schools and Colleges as Well as by the Working Entomologist
Packard, A. S. (Alpheus Spring)
Insects
On the other hand, we have been entirely mistaken in reference to
the mechanism which lowers the wings. The muscle concerned, that is
_kd_, is not at all the antagonist of the elevator muscle of the
wing, since it is placed close by this latter, but nearer to the
thoracic wall. But then, how does it come to be the counterpart of
its neighbor? In fact, the lever of the wing is situated in the
projecting piece alone. The extensor muscle of the wing does not
pull on the power arm, but on the resistant arm on the other side of
the fulcrum (_c_). The illustration shows, however, how such a case
is possible. The membrane of the joint fastening the wing-stalk to
the thorax is turned up outwards below the stalk like a pouch. The
tendon of the flexor of the wing passes through this pouch to its
point of attachment (_c_) lying on the other side of the fulcrum
(_d_). Thus it is very simply explained how two muscles which act in
the same direction can nevertheless have an entirely contrary
working power.
This is in a way the bare physical scheme of the flying machine by
the help of which we shall more easily become acquainted with its
further details.
Dragon-flies are unquestionably the most suitable objects for the
study of the muscles pulling directly on the wing itself. If the
lateral thoracic wall (Fig. 173) be removed or the thorax opened
lengthwise there appears a whole storehouse of muscular cords which
are spread out in an oblique direction between the base of the wing
and the side of the thoracic plate. There is first to be
ascertained, by the experiment of pulling the individual muscles in
the line with a pincers, which ones serve for the lifting and which
for the lowering of the wings. In dragon-flies the muscles are
arranged in two rows and in such a way that the flexors or
depressors (_s_, 1 bis) cling directly to the thoracic wall (compare
also the muscle _dk_ in Fig. 172 and _se_ in Fig. 174), while the
raiser or extensor (_h_ 1, to _h_ 2, Fig. 172, _hi_ and Fig. 174
_he_) lie farther in. The form of the wing-muscles is sometimes
cylindrical, sometimes like a prism, or even ribbon-like. However,
the contracted bundles of fibres do not come directly upon the
joint-process we have described, but pass over often indeed at a
very considerable distance from them, into peculiar chitinous
tendons. These have the form of a cap-like plate, often serrate on
the edge, which is prolonged into a thread, which should be
considered as the direct continuation of the base of the wings. The
wings, therefore, sink down into the thoracic cavity as if they were
a row of cords ending in handles where the strain of the muscles is
applied.
[Illustration:
Public-domain text, read in full here on John Shaqi.
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