A Text-book of Entomology: Including the Anatomy, Physiology, Embryology and Metamorphoses of Insects for Use in Agricultural and Technical Schools and Colleges as Well as by the Working EntomologistPackard, A. S. (Alpheus Spring)
Science
A Text-book of Entomology: Including the Anatomy, Physiology, Embryology and Metamorphoses of Insects for Use in Agricultural and Technical Schools and Colleges as Well as by the Working Entomologist
Packard, A. S. (Alpheus Spring)
Insects
“The hypopharynx, a continuation of the lower edge (_lamina_) of the
pharynx, most generally free, more or less produced, acute
anteriorly, forms with the labrum the tube of the pump (_antliæ_).
(The hypopharynx when obsolete, or coalesced with the canal of the
proboscis, is the _theca_; in such a case the siphon or tube is
formed by the theca and labrum.) Meanwhile the hypopharynx, the
largest of all the trophi (_omnium trophorum maximus_), constitutes
the chief piercing organ (_telum_) of Diptera. The hypopharynx is
moved by protractor, most generally quite or very powerful, and by
retractor muscles.
“The efferent duct of the thoracic salivary glands (_ductus
salivalis_) perforates the hypopharynx, more or less near the base,
that the saliva may be ejected through the canal into the wound, or
that it may be conducted along the labella. Very rarely the salivary
duct, perforating the hypopharynx, is continued in the shape of a
free, very slender tube.
“The salivary duct behind the base of the hypopharynx forms the
receptacle or _receptaculum_, provided with retractor and levator
muscles.”
[Illustration:
FIG. 83.—Labrum-epipharynx (_lbr_ and _eph_) and hypopharynx (_hyp_)
of _Tabanus brominus_: _oe_, posterior cylindrical portion of the
œsophagus; _a_, anterior swollen portion of the same; _ph_, pharynx;
_ph.m_, pharyngeal muscle; _p.ph_, protractor muscle of the pharynx;
_r.oe_, retractor muscle of the œsophagus; _r.ph_, retractor muscle
of the pharynx; _f.oe_, flexor muscle of the pharynx; _t.oe_,
twisting muscle of the œsophagus; _s.r_, receptacle of the salivary
duct; _l_, its elevator muscle; _s_, its retractor muscle; _cl_,
clypeus.—After Meinert.
]
[Illustration:
FIG. 84.—Œsophagus (_oe_), pharynx (_ph_) with epipharynx and labrum
(_lbr_) of _Asilus atricapillus_: _m_, _ph_, pharyngeal muscle;
_sr_, salivary receptacle; _t_, twisting; _r_, _l′r_, retractor
muscles; other lettering as in Fig. 83.—After Meinert.
]
It has been carefully studied by Meinert in a species of Culex (Fig.
81), Simulium (Fig. 82), Tabanus (Fig. 83), and in Asilus (Fig. 84),
where it is seen to attain enormous proportions. In the Hymenoptera,
this organ in its most specialized condition is a trough-like rod,
adapted for lapping nectar (Fig. 85, 86, _hyp_). The tongue or
hypopharynx of the honey-bee has been elaborately described by Cheshire
in his Bees and Bee Keeping.[18] He calls it the tongue or ligula. It is
situated in a tube formed by the maxillæ and labial palpi, and can be
partially retracted into the mentum. He states that it can move up and
down in the tube thus formed, and then describes it as covered by a
hairy sheath, its great elasticity being due to a rod running through
its centre enabling it to be used as a lapping tongue. The sheath
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