The most striking features of the =basal surface= are as follows: The
paramastoid processes are extremely long, less flattened than in the
horse and ox, and nearly vertical. At the inner side of the root of each
is the hypoglossal foramen, and in front of it are the stylo-mastoid
foramen and a deep cavity in which the hyoid process is concealed. The
bulla ossea is long, compressed laterally, and bears a sharp, short,
muscular process. The basioccipital is wide and flattened; it bears a
median crest and two lateral tubercles. The posterior nares are small
and are wider below than above. On either side of them is the tuberosity
of the palate bone, and above this is the pterygoid fossa. The palate is
remarkably long and is relatively narrow. It constitutes about
two-thirds of the entire length of the skull. It is widest between the
canines and premolars and narrow at each end. It is marked by a crest
medially and by the palatine foramen and groove laterally. The anterior
part bears transverse ridges. It is moderately arched from side to side.
In some specimens it is nearly straight or slightly concave in its
length; in others it curves upward to a variable degree in front. The
posterior end always slopes upward more or less.
The =posterior= or =nuchal surface= is remarkable for its height and the
breadth of the occipital crest. The central part above the foramen
magnum is smooth and concave from side to side, and is bounded laterally
by ridges, which converge below and end on two tubercles at the upper
margin of the foramen magnum. The surface is separated from the temporal
fossæ by the temporal crests, which curve downward and outward and blend
with the external auditory canals. The mastoid process has the form of a
plate which overlaps the root of the paramastoid process and bears a
crest on its anterior part.
The =cranial cavity= is small, in spite of the great size of the
cranium; the discrepancy is due to the enormous development of the
frontal sinuses in the adult. It is relatively longer, but much lower
than that of the ox. Its width is greatly diminished between the orbits.
The olfactory fossæ are extensive and very oblique. The floor resembles
that of the ox, but the foramen ovale is absent, the dorsum sellæ is
more developed, and the foramen lacerum basis cranii is like that of the
horse. Two oblique lateral crests clearly mark the limit between the
cerebral and cerebellar compartments. The tentorium osseum and the
parieto-temporal canals are absent.
The =nasal cavity= is very long. Its posterior part is divided by a
horizontal plate into olfactory and respiratory parts. The olfactory
part or fundus is above, and contains the ethmoturbinals and ethmoidal
meatuses. The lower part is continuous with the inferior meatus and
leads to the pharyngeal orifice; hence it is sometimes called the
naso-pharyngeal meatus. The bony roof is almost complete in front on
account of the great length of the nasal bones.
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