The =lateral cartilages= (Cartilagines ungulæ) are rhomboid curved
plates, which surmount the wings on either side. They are relatively
large and extend above the margin of the hoof sufficiently to be
distinctly palpable. The abaxial surface is convex, the axial concave.
The upper border is convex and thin; the lower is thicker and is in part
attached to the wing. The anterior end is attached by ligament to the
side of the second phalanx. The posterior end curves toward its fellow
at the heel, and is perforated by numerous foramina for the passage of
veins. The central part is mainly hyaline, the periphery mainly fibrous.
It will be noted that the size and form of the wings vary much in
different specimens. In the new-born foal the wing is a small, pointed
projection. Later the process of ossification invades the lower part
of the cartilage to a varying extent. In some cases the greater part
of the cartilage is ossified—a condition commonly termed “sidebone.”
=Development.=—The ossification of the terminal phalanx is peculiar.
While the proximal articular part is still cartilaginous, a
perichondrial cap of bone is formed in relation to the hoof. Later the
process extends into the upper part.
=Structure.=—The interior of this bone is channeled by numerous canals
for vessels, most of which radiate from the semilunar canal to the wall
surface; these are not canals for nutrient vessels of the bone, but
transmit arteries to the matrix of the hoof. Thick layers of compact
substance are found at the articular and flexor surfaces and the
extensor process, _i. e._, at the points of greatest pressure and
traction.
THE SESAMOID BONES
The two =proximal= or =great sesamoids= (Ossa sesamoidea phalangis
primæ) are situated behind the distal end of the large metacarpal bone,
and are closely attached to the first phalanx by strong ligaments. Each
has the form of a three-sided pyramid. The =anterior= or =articular
surface= conforms to the corresponding part of the distal end of the
large metacarpal bone. The =posterior= or =flexor surface= is flattened
and oblique; in the fresh state it is covered by a layer of cartilage
which also fills the interval between the opposed borders of the two
bones, and forms a smooth groove for the deep flexor tendon. The
=abaxial surface= is concave, and gives attachment to part of the
suspensory ligament; it is separated from the posterior surface by a
rough everted border. The =base= faces downward, and furnishes
attachment to the inferior sesamoidean ligaments. The =apex= is directed
upward and is rounded.
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