The external characters of an ordinary tapeworm may be gathered from
Figs. 135 and 136. A very small “head” continued behind into a small
“neck” can be distinguished, and a very large number of joints, of which
the first are very small and short, while those which follow get larger
and larger the further they are from the head. The tapeworm at first
consists only of a head which holds on to the lining of the intestine by
means of suckers or similar structures. The hinder part of the head next
elongates to form the so-called “neck,” and a cross-wall is quickly
developed a little way in front of the hind end of this neck. By this
division is formed the first joint, which, though forthwith tolerably
independent, remains attached to the neck of the tapeworm head, until a
new joint is separated off from the hinder part of the neck, immediately
in front of the first joint. Every new joint is similarly formed by the
constriction of the hinder part of the neck. Each joint possesses a
complete set of hermaphrodite reproductive organs; it becomes sexually
mature without reference to other joints, breaks off, and creeps about,
even for considerable distances, and finally ruptures, so that the
numerous eggs are liberated. Each joint may therefore be regarded as an
individual, though the tapeworm, considered as a whole, also possesses a
certain individuality. The tapeworm is a species of animal which exists
in two forms. One of these is the sexually reproducing joint, which
leaves the original host, and from the eggs of which are developed
larvæ, which become, in another host, _bladder-worms_, or _measles_.
These develop a _head_, which is the asexual second form of the
tapeworm. This, therefore, is a case of metagenesis (p. 16). It appears,
from what has been said, that the head can in no sense be compared to
the head of an ordinary animal; it has neither mouth nor jaws, for
tapeworms do not possess a gut. The head and joints take up osmotically
through their walls the nutritive matter found in the intestine of their
host. There are suckers and, in several species, a circlet of hooks on
the head (Fig. 136, _a_), by which the animal holds fast to the
intestinal wall. Thousands, or even tens of thousands, of eggs may be
produced by a single joint. The joint is liberated when the eggs begin
to ripen, often before. In some species several adjacent joints are
pinched off at the same time. The tapeworm joints leave the intestine of
their host either alone or with the dung. After falling to the ground
they creep along like snails, and attach themselves to various plants.
It is obvious that a tapeworm joint or a number of the eggs produced by
it may easily be taken up by a grazing animal. The eggs of a particular
kind of tapeworm are only able to develop further if they reach the
intestine of a particular kind of host. It is undoubtedly true that, as
a rule, only a relatively small number of the eggs produced by a
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