Among pseudopod forming organisms, the highest development of
contractility is found in the foraminifera. As is well known, these
organisms form finely anastomosing pseudopods which frequently cover the
substratum with a network of protoplasmic strands. The terminal sections
of these strands are frequently so thin and transparent that they cannot
be seen easily with the microscope. As a rule the granular endoplasm is
observable only in the main body of the organism and in the larger
trunks of the pseudopods. Much the larger part of the pseudopods, as
measured lineally, is devoid of granular endoplasm. The great power of
contractility and the speed with which contraction may occur in
_Biomyxa_, a fresh water foraminifer, have already been mentioned
(Figure 12, p. 47). Similar observations have been recorded by other
observers, recently by Schultz (’15), who compares the contractility of
foraminiferan pseudopods to that of rubber bands. In fact as one watches
the movements of a _Biomyxa_, for example, under moderately high
magnification, one gains the impression that there seems to be no
restriction imposed upon the extent of contractility in the pseudopods.
They seem to possess perfect elasticity. As to the transformation of
endoplasm into ectoplasm, little can be said, owing to the transparency
of the protoplasm. But the whole of the pseudopod, when forming, seems
to stream forward. As in _Difflugia_, the interior streams flow at about
the same rate as the pseudopod as a whole advances. The highly developed
power of contractility however demands rapid changes in phase of the
colloidal system, and also a thick consistency. The behavior of pieces
of the pseudopodial network, when cut from a _Biomyxa_, shows clearly
that the protoplasm is actually thick, as compared with that of an
_Amoeba proteus_. When a _Biomyxa_ is contracted into a spherical mass,
the interior exhibits continual rapidly streaming movements. Some of
these are rotational but most of them are radial. All of the streams
frequently change their direction and extent. No corresponding changes
are visible in the outer peripheral layer.
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