When a wave in the path for some reason becomes unusually long, there is
likely to be a very abrupt and decided change in the direction of
movement, which is away from the _convex_ side of the wave. Figure 37 is
inserted here to illustrate this point. The ameba should have turned to
the left at 3:43 to keep the waves of typical size, and at 3:45 a
pseudopod was extended in this direction a short distance, but again the
curve toward the right persisted. But at 3:48½ the ameba broke up into
several pseudopods at right angles with the main axis, and through one
[Illustration: Figure 37. An illustration showing that sudden changes
from the expected direction of the wavy path are centrifugal, not
centripetal; that is, away from the focus of the wave, not toward it.
The tendency to break away from the first smooth wave became apparent at
3:45, as indicated by the extension of pseudopods near the anterior end.
In the stage about two minutes later, a number of small pseudopods were
thrown out in various directions. At 3:48½ several large pseudopods were
thrown out near the anterior end almost at right angles to the main
axis, instead of at an angle of about sixty degrees or less, as is the
usual case. The same thing occurred at 3:57, except that the angle of
the pseudopod was not so large.]
of these the ameba moved on with the reestablishment of the wavy path.
The tendency to wave formation evidently has to overcome resistance of
some sort.
[Illustration: Figure 38. _Amoeba dubia_ usually moves with numerous
large pseudopods, but this illustration shows that there is very good
reason for concluding that there is a tendency in this ameba to move in
wavy paths. Length of the ameba, 400 microns.]
Although amebas in clavate shapes describe the smoothest waves in their
paths, waves may also be detected in the paths of amebas that habitually
form many pseudopods. The path of an _Amoeba dubia_ is shown in Figure
38. The ameba moved on an opal surface under light-controlled
conditions. If we had not already seen how _proteus_, _discoides_, and
especially _bigemma_ formed smooth waves in their paths, we should
hardly be able to understand the apparently aimless path of _dubia_. But
having seen how a regular succession of smooth waves appears under
favorable conditions in the paths of these amebas, there can be little
question but that the staggering path of a _dubia_ also is to be
interpreted as a succession of waves, although they are somewhat
irregular.
These four species of amebas, _proteus_, _dubia_, _discoides_ and
_bigemma_ are the only species that have been specially investigated as
to their paths, and they all show such paths, as we have seen. The
presumption is strong therefore that it is a common characteristic of
amebas.
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