Other specimens from Iowa are intergrades with _spadix_, or if not with
_spadix_, with the animal of northwestern Iowa which in some ways
combines the characters of _longicauda_ and _spadix_. For example, no.
2665, Coe College, an adult male from Davenport, has the anterior part
of the skull (all that is preserved) heavily ridged as in _spadix_ and
in addition, the underparts are marked with the shade of reddish
displayed by topotypes of _spadix_ and with some yellowish as seen in
_longicauda_. The color pattern, however, is as in _primulina_. A young
male, no. C-51, Iowa State College, from Kelley, Story County, has
anteriorly truncate bullae as are more frequently found in the
_longicauda-spadix_ stock of northwestern Iowa, than in _primulina_. In
other respects, the animal, in so far as can be judged from the broken
skull, agrees with _primulina_ as it certainly does in color, color
pattern, and external measurements. An adult male, no. 499a, Iowa State
College, from 2 miles east of Ledges St. Park, in Boone County, in
short body, size of teeth, and size of skull, in so far as the broken
parts can be measured, resembles _primulina_ more closely than it does
any other subspecies. The long tail, long hind foot, wide extent of the
light-colored underparts, and extension of the color of the underparts
onto the hind feet are more as in _spadix_. Other intergrades with
_spadix_ from Iowa are mentioned in the account of _spadix_.
The specimen from Swartz, Louisiana, suggests intergradation with
_arthuri_ in that the anteromedial part of the tympanic bulla is less
inflated than in typical _primulina_.
Intergrades with _longicauda_ are available from Riley and Pratt
counties, Kansas. No. 7182, Univ. Kans., subadult male in winter
pelage, from near Winkler, has a skull of larger size as in
_longicauda_ with which race it seems to agree in large size of body,
tail and hind foot, although the collector's measurements are lacking.
Color pattern and relative proportions of the skull throughout are as
in _primulina_. The young male, no. 3495, Univ. Kans., from Pratt,
Kansas, agrees in external measurements and large size of skull with
_longicauda_, but has the color and color pattern precisely as in
_primulina_. The teeth are smaller as in _primulina_. Immaturity
prevents judging of its relationships on the basis of relative
proportions of the skull.
Public-domain text, read in full here on John Shaqi.
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