The distinctness of _M. f. olivacea_ from _M. f. peninsulae_ is not
satisfactorily established due to inadequate material of _peninsulae_.
Differences shown by the specimens seen indicate that, as compared with
_olivacea_, _peninsulae_ is larger, has transversely wider
light-colored underparts which possess more yellow, and a larger skull
with more inflated tympanic bullae. In each of these characters,
_olivacea_ is intermediate between _noveboracensis_ on the north and
_peninsulae_ on the south. The question arises, therefore, whether the
animals here recognized under the name _olivacea_ really constitute a
recognizable subspecies or instead are only representatives of a
subspecies which reaches its extreme development in Florida. In the
latter event, the name _peninsulae_ would apply to all. Examination of
more material from Florida, especially from the southern half of
Florida, will be necessary to answer this question.
This large weasel of the southeastern United States is remarkably
different from _noveboracensis_. Indeed, were it not for actual
intergrades such as the two from Fort Payne, Alabama, and York, South
Carolina, which are described in the account of _M. f. noveboracensis_,
and the six specimens from northwestern Alabama, which are referred to
_olivacea_, the systematist, I believe, would have little or no
hesitancy in designating the two as distinct species, especially on the
basis of differences to be seen in the skull.
Not only are the two forms structurally more different than usually is
the case but between two geographically, adjacent subspecies of the
same species of mammal, but the belt where intergradation occurs
appears to be narrow. Nevertheless, when material of the two races is
laid out in geographic order, and examined in mass, certain features
are seen to undergo gradual change as a person's eye travels from
specimens from, say, the center of the range of _noveboracensis_ to
specimens from southern localities adjoining the territory occupied by
_olivacea_. One of these features subject to gradual change is the
color of the underparts. Beginning at the Adirondacks of New York
where a large number of the specimens have white underparts, the
underparts become more intensely yellowish southward through the range
of _noveboracensis_ into that of _olivacea_. Indeed, this progressive
trend seems to continue right on southward through the range of
_olivacea_ into that of _peninsulae_. Turning in the opposite direction
we find that the least width of the underparts decreases gradually
northward toward the range of _noveboracensis_. There is, likewise, a
decrease to the northward in length of the skull and relative, as well
as actual, narrowing of the braincase and tympanic bullae. However, in
least width of color of underparts and the mentioned cranial features,
the trend stops relatively abruptly at the southern boundary of the
geographic range of _noveboracensis_ and does not continue on,
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