_Mustela frenata spadix_ has just one structural feature of a "unique"
kind which serves to differentiate it from the geographically adjoining
subspecies. This feature is large size. The other diagnostic characters
ascribed to _spadix_ are of an intermediate sort--intermediate as
between two extremes, one found to the westward in _longicauda_ and the
other to the eastward in _noveboracensis_. For example, the
dark-colored upper parts are merely darker than in _longicauda_ and
merely lighter than in _noveboracensis_. The color is not "different";
it is only "intermediate." Furthermore, each of the characters ascribed
to _spadix_, including large size itself, undergoes change from one
part of its geographic range to another; the characters are not
constant over a wide area. Indeed, excepting the large size which
remains relatively uniform over the northern two-thirds of the range,
no two localities have been found from which the specimens can be said
really to agree in characters.
By way of illustration, the coloration of the upper parts may be cited.
Near the range of _noveboracensis_ the average coloration of
individuals from one locality is only a little lighter than in
_noveboracensis_. Farther westward the average coloration is a little
lighter and farther westward yet, toward the range of the extremely
light colored _longicauda_, the average coloration is lighter still.
Although all these animals are darker than _longicauda_ and lighter
than _noveboracensis_, those from the three places do not agree among
themselves. Because of the lack of more than one character of a
"unique" kind and because of the inconstancy, geographically, of other
characters, and for that matter, lack of constancy geographically in
combination of characters, the writer regards _spadix_ as a barely
recognizable subspecies.
Examination of the specimens of _spadix_ shows that the individual
variation in a single species is greater in a region of intergradation
than it is some distance inside the borders of the geographic range of
a well-marked subspecies. This is illustrated by three specimens of _M.
f. spadix_ in fresh summer pelage from the single locality, Elk River,
Minnesota. In these, the color of the upper parts varies from a little
darker than Cinnamon Brown to Vandyke Brown. At any one locality well
within the range of _longicauda_, or _noveboracensis_, there is nowhere
nearly so much variation in color, even in much larger series of
specimens.
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