Excepting the greater mastoidal breadth and generally larger size of
the skull, the cranial features distinguishing males of _spadix_ from
_longicauda_ are features in which _spadix_ shows approach to
_noveboracensis_. This is true, in _spadix_, of the relatively longer
(in comparison with _longicauda_) rostrum, relatively lesser zygomatic
breadth, relatively shallower braincase measured at the anterior end of
the basioccipital, and relatively deeper skull as measured at the
posterior borders of the last upper molars. This same approach to
_noveboracensis_ already has been pointed out with respect to color of
the upper parts and is evident also in the relative shortness of the
tail which averages only 51 per cent of the length of the head and body
rather than 55 per cent as in _longicauda_.
Because the _longicauda_ type of animal previously has been regarded as
specifically distinct from the _noveboracensis_ type of animal, comment
is offered below on selected specimens, referred to _spadix_, which are
regarded as intergrades with _noveboracensis_ or with other subspecies.
No. 8722, Univ. Wisconsin, adult male, in the white winter coat, from
north central Itasca County, Minnesota, obviously has characters of _M.
f. spadix_ or _longicauda_ that occur to the west and _M. f.
noveboracensis_ of the east. Selected outstanding characters of
_longicauda_ are its long tail, anteriorly truncate tympanic bullae
and large teeth. Characters indicating its affinities with
_noveboracensis_ are smaller size of skull, general narrowness of
skull, and relatively low tympanic bullae. The skull is intermediate as
regards several individual structural features. For example, although
long and narrow and in this feature more nearly approaching
_noveboracensis_, the skull is wider than usual in that subspecies and
thus approaches that of _longicauda_ or _spadix_. The hind foot, in the
dried state, measures 47 millimeters. This large hind foot, obviously
long tail (the specimen lacks external measurements), and anteriorly
truncate bullae constitute basis for here referring the specimen to
_spadix_. However, the seemingly small size of the body and the narrow
skull clearly show relationship to _noveboracensis_.
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