Thus, the Weismannian theory of evolution has entirely fallen to pieces
with the removal of its fundamental postulate—the absolute stability
of germ-plasm. It only remains to mention once more the effects of
this removal upon the other side of his system—viz., the companion
postulate of the uninterrupted continuity of germ-plasm, with its
superstructure in his theory of heredity.
Briefly, these effects are as follows:—
1. Germ-plasm ceases to be continuous in the sense of having borne a
perpetual record of congenital variations from the first origin of
sexual propagation.
2. On the contrary, as all such variations have been originated by the
direct action of external conditions, the continuity of germ-plasm in
this sense has been interrupted at the commencement of every inherited
change during the phylogeny of all plants and animals, unicellular as
well as multicellular.
3. But germ-plasm remains continuous in the restricted, though still
highly important sense, of being the sole repository of hereditary
characters of each successive generation, so that acquired characters
can never have been transmitted to progeny “representatively,” even
although they have frequently caused those “specialized” changes in the
structure of germ-plasm which, as we have seen, must certainly have
been of considerable importance in the history of organic evolution.
4. By surrendering his doctrine of the _absolute_ stability of
germ-plasm on the one hand, and of its _perpetual_[68] continuity on
the other, Weismann has greatly improved his theory of heredity. For,
whatever may be thought of his recent additions to this theory in
the way of elaborate speculation touching the ultimate mechanism of
heredity, it is a great gain to have freed his fundamental postulate
of the continuity of germ-plasm from the two further postulates which
have just been mentioned, and the sole purpose of which was to provide
a basis for his untenable theory of evolution.
5. In my opinion it only remains for him to withdraw the last remnant
of his theory of evolution by cancelling his modified and even less
tenable views on amphimixis, in order to give us a theory of heredity
which is at once logically intact and biologically probable.
6. The theory of germ-plasm would then resemble that of stirp in all
points of fundamental importance, save that while the latter leaves
the question open as to whether acquired characters are ever inherited
in any degree, the former would dogmatically close it, chiefly on the
grounds which I have considered in Appendix II. It seems to me that in
the present state of our knowledge it is more prudent to follow Galton
in suspending our judgement with regard to this question, until time
shall have been allowed for answering it by the inductive methods of
observation and experiment.
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