Here, then, we have actual proof of “hereditary individual variations”
among a-sexually propagating organisms, sufficient in amount to
have given origin, not merely to “individual differences,” but to
innumerable species, and even genera. Consequently Weismann allows
that the criticism abolishes this line of evidence in favour of the
absolute stability of germ-plasm[26]. Consquently, also, we must now
add, in whatever measure the alleged fact would have corroborated
the theory had it been proved to be a fact, in that measure is the
theory discredited by proof that it is not a fact. For, if the theory
were sound, this particular fact would certainly have admitted of
demonstration: therefore the proof that it is not a fact—but the
reverse of a fact—amounts at the same time to a disproof of the
theory[27].
The only other line of evidence to be adduced in favour of the absolute
stability of germ-plasm is that which is furnished by the high
antiquity of some specific types, by the facts of atavism, and by the
persistency of vestigial organs. But this line of evidence is as futile
as the other. Nobody has ever questioned that hereditary characters
are persistently stable as long as they are persistently maintained by
natural selection; and this, according to Weismann himself, must have
been the case with all long-enduring species: these, therefore, fail to
furnish any evidence of the _inherent_ stability of germ-plasm, which
is the only point in question.
Again, as regards the facts of atavism, nobody is disputing these
facts. What we are disputing is whether the _degree_ of inherent
stability which they unquestionably prove can be rationally regarded
as such that it may endure, not merely for such a comparatively small
number of generations as these facts imply, but actually for any
number of generations, or through the practically infinite series of
generations that now intervene between the higher metazoa and their
primeval parentage in the protozoa. Clearly, the ratio between these
two things is such that no argument derived from the facts of atavism
can be of any avail for the purposes of this Weismannian doctrine.
Lastly, as regards vestigial organs, the consideration that,
surprisingly persistent as they unquestionably are, nevertheless they
do eventually disappear, seems to prove that the power of heredity
does in time become exhausted, even in cases most favourable to
its continuance. That it should thus become finally exhausted is
no more than Darwin’s theory of perishable gemmules, or Galton’s
theory of a not absolutely stable stirp, would expect. But the fact
is irreconcilable with Weismann’s theory of an absolutely stable
germ-plasm.
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