It does not appear to me that very much has to be said, either for
or against the former postulate, on merely antecedent grounds, or
grounds of general reasoning. Therefore I relegated to an Appendix
my examination of what Weismann has argued on these grounds, while
in the text I considered only what he has advanced as evidence _a
posteriori_. Here, as we saw, he has developed three distinct lines
of verification—viz. (A) the migration of germ-cells in some of
the _Hydromedusae_,(B) the early separation of germ-cells in the
ontogeny of certain Invertebrata, and (C) the alleged invariability of
organisms which are produced parthenogenetically. But we have seen,
with respect to (A), that the specialized character of germinal cells
is a fact which every theory of heredity must more or less recognize;
and, therefore, that the migration of these cells, wherever it may be
found to occur, does not lend any peculiar countenance to Weismann’s
theory. There may be many reasons for such migration other than the one
which this theory assigns; while the reason which it does assign is
rendered improbable by the consideration that in the _Hydromedusae_
the material of heredity is already and richly diffused throughout the
general tissues. (B) and (C) are both contrary to fact; and, therefore,
in whatever measure they would have corroborated the theory had they
proved to be true, in that measure must they be held to discountenance
the theory now that they have been shown to be false.
It appears, then, that there is no evidence in support of the postulate
of the _perpetual_ continuity of germ-plasm. There is nothing to show
the necessary non-inheritance of acquired characters. The only evidence
which one can recognize as good, is that which makes equally in
favour of the theory of stirp—or rather, of the well-known fact that
congenital characters are at any rate much _more_ heritable than are
acquired: which, it is needless to repeat, is a widely different thing
from proving—or even rendering probable—the _absolute_ restriction of
germ-plasm to a separate “sphere” of its own “since the origin of life.”
Public-domain text, read in full here on John Shaqi.
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