First of all, Weismann has now profoundly modified his theory of polar
bodies. For, owing to certain more recent researches of Professor O.
Hertwig, he very candidly allows:—“My previous interpretation of the
first polar body as the removal of ovogenetic nucleo-plasm from the egg
must fall to the ground: about this there is no possible doubt[36].”
He now regards both polar bodies as concerned in the same function of
removing superfluous germ-plasm. Therefore one-half of his previous
theory is abandoned: “the ovogenetic idio-plasm” is now supposed to be
simply absorbed in the course of ontogeny, as I had suggested in one
of the preceding chapters (pp. 42-46). The consequence is that he has
now nothing to oppose to the view which is likewise there suggested
(pp. 43-44)—viz., that his whole theory of polar bodies is rendered
needless and improbable by the fact that the very mode in which ova
are produced renders ample provision for the removal of any amount of
superfluous germ-plasm which the theory of germ-plasm may require.
It is needless to say, after what has already been said in the pages
just referred to, that in my opinion Professor Weismann has improved
his main system of theories by dropping this part of his subordinate
and, for the most part, separate theory of polar bodies. I only wish he
could have seen his way to dropping the whole.
* * * * *
Again, he has now fully considered the phenomena of repair,
regeneration, reproduction from somatic tissues, budding, and
graft-hybridization.
Touching the four former he takes the view which I have supposed that
he would (p. 53). As regards the latter, he fully accepts the fact of
an occasional transmission of characters from one species or variety of
plant to another by mere grafting[37]. But, although the explanation
which he gives of this fact may pass muster so far as the only case
which he deals with in detail is concerned, I do not see how it can do
so to many others. For the case which he considers is that of _Cystisus
adami_, where a bud of one species of Laburnum having been inserted
in the wood of another produced a shoot which presented intermediate
characters; and these have ever since been propagated by cuttings.
Weismann’s interpretation of the facts here is, “that they were due to
an abnormal kind of amphimixis, so that the idants of both species were
combined in the apical cell of the first shoot[38].” Now, although this
explanation may well apply to a case of graft-hybridization by means
of buds, it obviously cannot do so to any case where hybridization is
produced by the grafting of woody tissues. For here there is no “apical
cell” in the question; and therefore the difficulties which I have
adduced on page 82 remain. Possibly Weismann may dispute the fact of
hybridization in any of these cases; but, as he has not expressly done
so, I will not go into the question of evidence[39].
* * * * *
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