As such eminent botanists as Focke, and more recently De Vries, have
expressed much doubt with regard to these observations—or rather
interpretations—we must wait until these cases have been critically
re-investigated before attempting to account for them theoretically.
The chief difficulty we should meet with in any such explanation would
be due to the fact that we are here concerned with the influence of
the _germ-plasm_ of the sperm-cell on a tissue of another plant which
only constitutes _a part_ of this plant. It would thus be necessary to
assume that all the determinants of this germ-plasm are not active,
and that only those take effect which determine the nature of the
fruit.
Now, it does not appear that De Vries has looked into the matter on
his own account, as he merely refers to what Focke has said. And this
amounts merely to showing the dubious character of some half-dozen
cases which Focke gives as those which alone have fallen within his
cognizance. Why he does not mention any of the numerous cases which are
quoted by Darwin, I do not understand. Nor can I understand why he does
not consider what seem to be the particularly conclusive facts given on
p. 80,—i. e., where xenia appears to constitute “a needful preliminary
to fertilization.” But the whole matter is one for botanists to deal
with, and if any doubt attaches to it, at least the grounds of such
doubt should be fully stated. Still more, in my opinion, should the
matter be freed from any such doubt. The question—if there be a
question—is one of great interest from a merely physiological point
of view, while in relation to the fundamental problems of heredity
its importance is immense. Surely, then, any competent botanist who
disputes the facts ought to test them by way of experiment.
Public-domain text, read in full here on John Shaqi.
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