An Introduction to Nature-studyStenhouse, E. (Ernest)
History
An Introduction to Nature-study
Stenhouse, E. (Ernest)
Natural history -- Study and teaching; Nature study
The whole group of flowers borne by any one haulm is generally called
the ear or panicle. The ear in its turn consists of
several bodies called spikelets. The appearance of the ear varies
greatly in different grasses (Figs. 102 to 110) according as the stalks
of the spikelets are long and spread outwards from the haulm, as in the
meadow-grasses (Fig. 103), oats
(Figs. 99 and 108),
fescues (Fig. 102),
and others; short, as in the foxtails (Figs. 104 and 105),
timothy (Fig. 106), and sweet vernal grass
(Fig. 110); or absent altogether, as
in the wheat, barley, rye, rye grasses (Fig. 109),
and couch grass.
A single spikelet of a grass is shown diagrammatically in Fig. 100.
At the bottom are two boat-shaped bracts called glumes
(g, g), which almost or entirely cover the spikelet before it
opens. When the glumes have been removed a few flowers remain. Each
one is protected by two leaves called pales: outer (p₁)
and inner (p₂) respectively. The flower (B), as a rule, has
three stamens, the anthers being borne on long filaments and dangling
out to the wind (Fig. 101), and a pistil with two branching feathery
stigmas. In many grasses the outer pale bears a bristle called an
awn, well seen in “bearded” wheat, barley, and upright brome grass.
Fig. 101, A, represents a single spikelet of meadow fescue, with two
[Pg 131]
open flowers. The outer pales of this grass are not awned. Fig. 101,
B, shows a single flower of the same grass from which the outer pale
has been removed. The two little scales seen in front of the ovary (and
at e in Fig. 100) possibly represent all that
is left of a perianth (p. 121).
Fertilisation.—By the time the flowers have opened, the haulm
has usually grown so tall that they are lifted well above the leaves.
The stamens hang their anthers loosely out to the wind, and, as the
slender haulm sways in the breeze, the pollen is readily detached and
carried to other flowers, perhaps miles away.
It is in order to catch the wind-borne pollen-grains that the stigmas
of grasses are branched and feathery. A great waste of pollen is
obviously entailed by this method. But on the other hand it must
be remembered that insect-pollinated flowers have to pay for their
privileges by storing nectar in cunningly hidden pockets, and by
advertisement-expenses, all of which the grasses avoid.
Fig. 101.—Meadow Fescue.
A, spikelet with two open flowers (× 1½);
B, a flower from which the outer pale has
been removed (× 6).
Public-domain text, read in full here on John Shaqi.
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