An introduction to the study of fishesGünther, Albert C. L. G. (Albert Carl Ludwig Gotthilf)
Science
An introduction to the study of fishes
Günther, Albert C. L. G. (Albert Carl Ludwig Gotthilf)
Fishes; Ichthyology
As far as the observations go at present, no distinct bathymetrical
regions, which would be characterised by peculiar forms, can be
defined. The depths from 200 to 600 fathoms are inhabited by
numerous forms, still strongly reminding us of surface types. To this
fauna belong the few Chondropterygians of the deep sea, a Sebastes
and Setarches, a Beryx and Polymixia, a Cottus, etc.; but they are
associated with many others which descend to the greatest depths.
And before anything like a division into bathymetrical zones can be
attempted, the observations of the “Challenger” expedition must be
confirmed and supplemented by other series of similar systematic
observations. One of the most startling conclusions at which we would
have to arrive from the “Challenger” observations is, that some of the
species of Deep-sea fishes would range from a depth of some 300 fathoms
down to one of 2000 fathoms; or, in other words, that a fish which
has once attained in its organisation to that modification by which
it is enabled to exist under the pressure of half a ton, can easily
accommodate itself to one of two tons or more,--a conclusion which is
not in accordance with anatomical facts, and which must be confirmed by
other observations before we can adopt it. But if the vertical range of
Deep-sea fishes is actually as it appears from the “Challenger” lists,
then there is no more distinct vertical than horizontal distribution of
Deep-sea fishes.
The greatest depth reached hitherto by a dredge in which fishes were
enclosed is 2900 fathoms. But the specimens thus obtained belong to a
species (_Gonostoma microdon_), which seems to be extremely abundant
in upper strata of the Atlantic and Pacific, and were therefore most
likely caught by the dredge in its ascent. The next greatest depth,
viz., 2750 fathoms, must be accepted as one at which fishes undoubtedly
do live; the fish obtained from this depth of the Atlantic, _Bathyophis
ferox_, showing by its whole habit that it is a form living on the
bottom of the ocean.
The fish-fauna of the deep sea is composed chiefly of forms or
modifications of forms which we find represented at the surface in the
cold and temperate zones, or which appear as nocturnal pelagic forms.
The Chondropterygians are few in number, not descending to a depth of
more than 600 fathoms. The Acanthopterygians, which form the majority
of the coast and surface faunas, are also scantily represented; genera
identical with surface types are confined to the same inconsiderable
depths as the Chondropterygians, whilst those Acanthopterygians which
are so much specialised for the life in the deep sea as to deserve
generic separation, range from 200 to 2400 fathoms. Three distinct
families of Acanthopterygians belong to the deep-sea fauna, viz.
_Trachypteridæ_, _Lophotidæ_, and _Notacanthidæ_; they respectively
consist of three, one, or two genera only.
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