An introduction to the study of fishesGünther, Albert C. L. G. (Albert Carl Ludwig Gotthilf)
Science
An introduction to the study of fishes
Günther, Albert C. L. G. (Albert Carl Ludwig Gotthilf)
Fishes; Ichthyology
The form of the _bulbus_ (Fig. 48) is subhemispherical, the cornea
(_co_) being flat. If it were convex, as in higher Vertebrates, it
would be more liable to injury; but being level with the side of the
head the chances of injury by friction are diminished. The _sclerotica_
(_sc_) is cartilaginous in Chondropterygians and Acipensers, fibrous
and of varying thickness in Teleosteans, in the majority of which it
is supported by a pair of cartilaginous or ossified hemispheroid cups
(_c_). In a few fishes, as in _Ceratodus_, _Xiphias_, the cups are
confluent into one cup, which possesses a foramen behind to allow the
passage of the optic nerve (_o_). The _cornea_ of _Anableps_ shows an
unique peculiarity. It is crossed by a dark horizontal stripe of the
conjunctiva, dividing it into an upper and lower portion; also the iris
is perforated by two pupils. This fish is observed to swim frequently
with half of its head out of the water, and it is a fact that it can
see out of the water as well as in it.
The membranes situated between the sclerotica and retina are
collectively called _choroidea_, and three in number. The one in
immediate contact with the sclerotic, and continued upon the iris,
is by no means constantly present; it is the _membrana argentea_
(_a_), and composed of microscopical crystals reflecting a silvery or
sometimes golden lustre. The middle layer is the _membrana vasculosa_
s. _halleri_ (_v_), the chief seat of the ramifications of the choroid
vessels; the innermost layer is the _membrana ruyscheana_ or _uvea_
(_u_), which is composed of hexagonal pigment-cells, usually of a deep
brown or black colour.
In many _Teleostei_ a _rete mirabile_ surrounds the entry of the optic
nerve; it is situated between the membrana argentea and vasculosa, and
called the _choroid gland_ (_ch_). It receives its arterial blood from
the artery issuing from the pseudobranchia; the presence of a choroid
gland always being combined with that of a pseudobranchia. Teleosteans
without pseudobranchia lack a choroid gland. In the Palæichthyes, on
the other hand, the pseudobranchia is present and a choroid gland
absent.
The _iris_ (_i_) is merely the continuation of the choroid membrane;
its capability of contracting and expanding is much more limited than
in higher Vertebrates. The _pupil_ is generally round, sometimes
horizontally or vertically elliptical, sometimes fringed. In the
Rays and Pleuronectidæ a lobe descends from the upper margin of the
pupil, and the outer integument overlying this lobe is coloured and
non-transparent; a structure evidently preventing light from entering
the eye from above.
In most _Teleostei_ a fold of the _Choroidea_, called the _Processus
falciformis_ (_f_), extends from the vicinity of the entrance of the
optic nerve to the lens. It seems to be constantly absent in Ganoids.
Public-domain text, read in full here on John Shaqi.
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