An introduction to the study of mammals living and extinctLydekker, Richard
Science
An introduction to the study of mammals living and extinct
Lydekker, Richard
Mammals
Another equally well-marked and equally isolated, though far more
numerously represented and diversified order, is that of the Cetacea,
placed simply for convenience next to the Sirenia; with which, except in
their fish-like adaptation to aquatic life, they have little in common.
The old association of these orders in one group can only be maintained
either in ignorance of their structure or in an avowedly artificial
system. Among the existing members of the order, there are two very
distinct types, the toothed Whales or Odontoceti, and the Baleen Whales
or Mystacoceti, which present as many marked distinguishing structural
characters as are found between many other divisions of the Mammalia
usually reckoned as orders. Since the extinct Zeuglodonts, so far as
their characters are known, do not fall into either of these groups, but
are in some respects annectant forms, we have placed them provisionally,
at least, in a third group by themselves, named Archæoceti. There is
nothing known at present to connect the Cetacea with any other order of
Mammals; but it is quite as likely that they are offsets of a primitive
Ungulate as of a Carnivorous type, or perhaps of a still more generalised
mammalian stock.
The remaining Eutherian mammals are clearly united by the characters
of their teeth, being all heterodont and diphyodont, with their dental
system reducible to a common formula.
Although older views of the relationship of Ungulate mammals expressed
by the terms _Pachydermata_, _Ruminantia_, and so forth, still
linger in some corners of zoological literature, no single point in
zoological classification can be considered so firmly established as
the distinction between the Perissodactyle and Artiodactyle Ungulates;
both being in the existing fauna of the world perfectly natural and
distinctly circumscribed groups. The breaking-up of the latter into
four equivalent sections, the Pecora, Tylopoda, Tragulina, and Suina,
is equally in accordance with all known facts. Less certain, however,
is the association of the Proboscidea and the Hyracoidea with the true
Ungulates. By many zoologists they are each, although containing so very
few existing species, made into distinct orders; and much is to be said
in favour of this view. The discovery, however, of a vast number of
extinct species of Ungulates which cannot be brought under the definition
of either Perissodactyla or Artiodactyla, and yet are evidently allied
to both, and to a certain extent bridge over the interval between them
and the isolated groups just mentioned, make it necessary either to
introduce a number of new and ill-defined ordinal divisions, or so to
widen the scope of the original order as to embrace them all, considering
the Elephants and the Hyraces as representing suborders equivalent to
the great Perissodactyle and Artiodactyle groups. It is the latter
alternative that we have adopted.
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