An introduction to the study of mammals living and extinctLydekker, Richard
Science
An introduction to the study of mammals living and extinct
Lydekker, Richard
Mammals
_Multituberculata._—The name Multituberculata has been proposed for
the group exhibiting the type of dentition last mentioned, and is
generally adopted, although the term Allotheria has been also suggested.
The essential characteristic of the dentition of this group is the
presence of a single scalpriform incisor on each side of the lower
jaw (Fig. 25) and of one larger incisor, and in some instances of one
or two smaller ones in each premaxilla (Fig. 24). These incisors are
separated by an interval or diastema from the first of the premolars.
The true molars, and in some instances the premolars (Fig. 24), are
characterised by having longitudinal rows of tubercles separated by
one or more grooves; there being either two or three of these rows in
the upper molars of those forms in which these teeth are known, while
there are, at least usually, only two in those of the lower jaw. In
other cases the premolars are of a secant type, with a highly convex
cutting-edge, and usually either serrated or obliquely grooved (Figs.
25, 26). From a certain resemblance between these secant premolars and
those of some of the smaller _Macropodidæ_ it was at one time considered
that we had in these mammals representatives of Diprotodont Marsupials.
The great difference in the structure of the molar teeth of these forms,
coupled with the circumstance that when the number of upper incisors is
reduced below three it is the second in place of the first which becomes
enlarged and opposed to the incisor of the lower jaw, seems to prevent
the acceptation of this view. Moreover, in their peculiar structure
the molars seem, on the whole, to make a nearer approximation to the
teeth of _Ornithorhynchus_ than to any other known mammal; and it has
accordingly been suggested that the Multituberculata may really represent
an order of Prototheria. Some support is afforded to this suggestion by
certain fragmentary bones from the Cretaceous of the United States, which
are regarded by Marsh as parts of a coracoid and interclavicle. The
peculiar character of the whole dentition of these forms indicates that
if they are really Prototherians they cannot be regarded as primitive and
ancestral types.
[Illustration: FIG. 25.—The right ramus of the mandible of _Plagiaulax
beklesi_; from the Purbeck of Swanage. Twice natural size. _i_, Incisor;
_m_, molar; _b_, coronoid process; _c_, condyle. (After Owen.)]
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