An introduction to the study of mammals living and extinctLydekker, Richard
Science
An introduction to the study of mammals living and extinct
Lydekker, Richard
Mammals
The scapula differs from that of all other mammals in that the ridge
corresponding to the spine of other forms is situated on the anterior
border instead of in the middle of the outer or dorsal surface. The
humerus is much expanded at its two extremities, and has a very prominent
deltoid crest, and a well-marked entepicondylar foramen.
The dorso-thoracic vertebræ are nineteen in number, and have no terminal
epiphyses to their bodies. The transverse processes of the cervical
vertebræ are of autogenous formation, and remain suturally connected with
the remainder of the vertebra until the animal is full-grown. Though in
this respect they present an approximation to the Sauropsida (Reptiles
and Birds), they differ from these classes, inasmuch as there is not a
gradual transition from these autogenous transverse processes of the neck
(or cervical ribs, as they may be considered) into the thoracic ribs, for
in the seventh vertebra the costal element is much smaller than in the
others, indicative of a very marked separation of neck from thorax, not
seen in the existing Sauropsida. The upper ends of the ribs are attached
to the sides of the bodies of the dorsal vertebræ only, and not to the
transverse processes. The sternal ribs are well ossified, and there are
distinct partly ossified intermediate ribs. The cerebral cavity, unlike
that of the lower Marsupials or the Reptiles, is large and hemispherical,
flattened below and arched above, and about as broad as long. The
cribriform plate of the ethmoid is nearly horizontal. The cranial walls
are very thin, and smoothly rounded externally, and the sutures become
completely obliterated in adult skulls, as in Birds. The broad occipital
region slopes upwards and forwards, and the face is produced into a long
and depressed rostrum. The bony palate is prolonged backwards, so that
the posterior nares are nearly on a level with the glenoid fossæ. The
mandible is without distinct ascending ramus; the coronoid process and
angle are rudimentary, and the two halves are loosely connected at the
symphysis. The fibula has a broad, flattened process, projecting upwards
from its upper extremity above the articulation, like an olecranon. In
the male there is an additional, flat, curved ossicle on the hinder and
tibial side of the plantar aspect of the tarsus, articulating chiefly
to the tibia, which supports in the adult a sharp-pointed perforated
horny spur, with which is connected the duct of a gland situated
beneath the skin of the back of the thigh, the function of which is
not yet clearly understood. (A rudimentary spur is found in the young
female _Ornithorhynchus_, but this disappears when the animal becomes
adult.) The stomach is subglobular and simple; the alimentary canal
has no ileo-cæcal valve, or marked distinction between large and small
intestine, but has a small, slender vermiform cæcum with glandular walls.
The liver is divided into the usual number of lobes characteristic of the
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