An introduction to the study of mammals living and extinctLydekker, Richard
Science
An introduction to the study of mammals living and extinct
Lydekker, Richard
Mammals
the actual progenitors of the Mammalia, and that they should rather be
regarded as a divergent branch of the stem which had given origin to the
Mammalia. Since that date observations made on the structure of the South
African Anomodonts have shown such an intimate connection between that
group and the Labyrinthodont Amphibians, that there can be no hesitation
in regarding the one as the direct descendant of the other; and we may
probably regard the Mammalia as having originated from the same ancestral
stock at the time the Amphibian type was passing into the Reptilian. From
this point of view, some of the mammalian features found in the more
specialised Anomodonts may probably be regarded as having been acquired
during a parallel line of development.
Both the Anomodontia and the Mammalia differ from the Amphibians in the
loss of the splint-like parasphenoid which underlies the basisphenoid
axis of the skull, and by the ossification of that axis; but while the
former have become monocondylic by the participation of the basioccipital
in the support of the cranium, the latter retain the Amphibian dicondylic
plan. The skull of the Anomodonts presents mammalian resemblances not
found in any other Reptiles, this being especially noticeable in the
region of the squamosal; and it is only in this group and mammals that
the temporal or zygomatic arch is a squamoso-maxillary one (see p.
37). The resemblance between the pectoral and pelvic girdles of the
Anomodonts and those of the Monotreme Mammals is noticed under the head
of the latter, where reference is also made to the similarity in the
structure of the humerus in the two groups. The pes of the Amphibia and
Anomodontia agree in having a distinct intermedium, tibiale, fibulare,
and centrale, whereas in other Reptiles these bones are not generally
distinct; in Mammals the intermedium, fibulare, and centrale are
distinct, and according to Cope’s interpretation there may be a distinct
tibiale.
_Classification._—In the present condition of the world, mammals
have become so broken up into distinct groups by the extinction of
intermediate forms, that a systematic classification is perfectly
practicable. Most of the associations of species, which we call “orders,”
and even the “suborders” and “families,” are natural groups. In
isolating, defining, and naming them, we are really dealing with facts
of nature of a totally different order from the artificial and fanciful
divisions formed in the infancy of zoological science.
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