Ancient Plants: Being a Simple Account of the past Vegetation of the Earth and of the Recent Important Discoveries Made in This Realm of NatureStopes, Marie Carmichael
Science
Ancient Plants: Being a Simple Account of the past Vegetation of the Earth and of the Recent Important Discoveries Made in This Realm of Nature
Stopes, Marie Carmichael
Paleobotany
On the other hand, there are not wanting scientists who consider the
_Abietineæ_ the living representatives of the most primitive and ancient
stock, though on the whole the evidence seems to indicate more clearly
that the Pine-tree group is specialized and highly modified. Their double
series of foliage leaves, their complex cones (whose structures are not
yet fully understood), and their wood all support the latter view.
Some, again, consider the _Taxeæ_ as a very primitive group, and would
place them near the Cordaiteæ, with which they may be related. Their
fleshy seeds, growing not in cones but on short special axes, support
this view, and it is certainly true that in many ways the large seeds,
with their succulent coats and big endosperm, are much like those of the
lower Gymnosperms and of several fossil types. Those, however, who hold
to the view that the Abietineæ are primitive, see in the _Taxeæ_ the
latest and most modified type of Gymnosperm.
It will be seen from this that there is no lack of variety regarding the
interpretation of Gymnosperm structures.
The Gymnosperms do not stand in such an isolated position as do the
Angiosperms. Whatever the variety of views held about the details of the
relative placing of the families within the group, all agree in
recognizing the evidence which enables us to trace with confidence the
connection between the lower Gymnosperms and the families of ferns. There
are many indications of the intimate connection between higher and lower
Gymnosperms. Between the series exist what might be described as
different degrees of cousinship, and in the lower groups lie unmistakable
clues to their connection with more ancient groups in the past which
bridge over the gaps between them and the ferns.
For the present, however, let us confine ourselves to the history of the
more important Gymnosperms, the discussion of their origin and the groups
from which they may have arisen must be postponed until the necessary
details about those groups have been mentioned.
To a consideration of the living families of _Araucareæ_, _Abietineæ_,
_Cupresseæ_, and _Taxeæ_ we can allow but a short space; their general
characters and appearance are likely to be known to the reader, and their
details can be studied from living specimens if they are not. For
purposes of comparison with the fossils, however, it will be necessary to
mention a few of the principal features which are of special importance
in discussing phylogeny.
The Araucariaceæ are woody trees which attain a considerable size, with
broad-based, large leaves attached directly to the stem. In the leaves
are a series of numerous parallel vascular bundles. The wood cells in
microscopic section show two rows or more of round bordered pits. The
cones are very large, but the male and female are different in size and
organization. The female cone is composed of series of simple scales
arranged spirally round the axis, and each scale bears a single seed and
a small ligule.
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