Ancient Plants: Being a Simple Account of the past Vegetation of the Earth and of the Recent Important Discoveries Made in This Realm of NatureStopes, Marie Carmichael
Science
Ancient Plants: Being a Simple Account of the past Vegetation of the Earth and of the Recent Important Discoveries Made in This Realm of Nature
Stopes, Marie Carmichael
Paleobotany
_Poroxylon_ is not common, and until recently had not been found in the
Lower Coal Measures of England. The plants appear to have been much
smaller than _Cordaites_, with delicate stems which bore relatively large
simple leaves. The anatomy of the root was that common in Gymnosperms,
but the stem had a very large pith, and the leaves were much like those
of _Cordaites_ in having parallel veins. An important character in the
anatomy of the stem was the presence of what is known as _centripetal
wood_. This must be shortly explained. In all the stems hitherto
considered, the first-formed wood cells (protoxylems, see p. 57)
developed at the central point of the wood, towards the pith (see fig.
19, _px_, p. 49). This is characteristic of all Angiosperms and the
higher Gymnosperms (except in a couple of recently investigated Pines),
but among the lower plants we find that part of the later wood develops
to the inner side of these protoxylem masses. The distinction is shown in
fig. 65.
[Illustration: Fig. 65.—A, Normal bundle of higher plant; _x_,
protoxylem on inner side next the pith _p_, and the older wood _w_
outside it, _centrifugal_ wood. B, Bundle with wood cells _c_ developed
on inner side of protoxylem, _centripetal_ wood; the arrow indicates
the direction of the centre of the stem.]
This point is one to which botanists have given much attention, and on
which they have laid much weight in considering the affinities of the
lower Gymnosperms and the intermediate groups between them and the ferns,
which are found among the fossils. In _Cordaites_ this point of
connection with the lower types is not seen, but in _Poroxylon_, which
has otherwise a stem anatomy very similar to _Cordaites_, we find groups
of _centripetal_ wood developed inside the protoxylem of primary bundles.
For this reason, principally, is _Poroxylon_ of interest at present, as
in its stem anatomy it seems to connect the _Cordaites_ type with that of
the group below it in general organization.
Ginkgoales.—Reference to p. 44 shows that _Ginkgo_, the Maidenhair tree,
belongs to the Ginkgoales, a group taking equal rank with the large and
complex series of the Coniferales. The Ginkgoales of the present day,
however, have but one living representative. _Ginkgo_ stands alone, the
single living species of its genus, representing a family so different
from any other living family that it forms a prime group by itself.
Had the tree not been held sacred in China and Japan, it is probable that
it would long since have been extinct, for it is now known only in
cultivation. It is indeed a relic from the past which has been
fortunately preserved alive for our examination. It belongs to the fossil
world, as a belated November rose belongs to the summer.
Public-domain text, read in full here on John Shaqi.
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